Ivan Arismendi

Assistant Professor
DEPT. of fisheries and wildlife


I am a quantitative aquatic ecologist interested in the links among water, land, and people. I focus on freshwater-terrestrial links, freshwater-marine links, invasive species impacts on aquatic and riparian ecosystems, and the consequences of climate change on freshwater food webs and ecosystems. I am also interested in issues related to diversity and inclusion in science.

Selected publications

Murphy CA, Arismendi I, Taylor GA, Johnson SL (2019) Evidence for lasting alterations to aquatic food webs with short-duration reservoir draining. PLoS ONE 14(2): e0211870. https://doi.org/10.1371/journal.pone.0211870

Flitcroft R, S Lewis, I Arismendi, R LovellFord, M Santelmann, M Safeeq, G Grant. 2016. Linking hydroclimate to fish phenology and habitat use with ichthyographs. PLoS ONE 11(12): e0168831. doi:10.1371/journal.pone.0168831

Arismendi I, BE Penaluna. 2016. Examining diversity inequities in fisheries science: A call to action. Bioscience 66(7):584-591.  Highlighted as the front-matter of the July issue

Vargas PV, I Arismendi, D Gomez-Uchida 2015. Evaluating taxonomic homogenization of freshwater fish assemblages in Chile. Revista Chilena de Historia Natural 88(16) DOI: 10.1186/s40693-015-0046-2

Arismendi I, B Penaluna, JB Dunham, C García de Leaniz, D Soto, I Fleming, D Gomez-Uchida, G Gajardo, P Vargas, J León-Muñoz. 2014. Differential invasion success of salmonids in southern Chile: patterns and hypotheses. Reviews in Fish Biology and Fisheries 24:919-941.

Marr SM, MP Marchetti, JD Olden, E García-Berthou, DL Morgan, I Arismendi, JA Day, CL Griffiths & PH Skelton 2010. Freshwater fish introductions in Mediterranean-climate regions: are there commonalities in the conservation problem? Diversity and Distributions 16(4):606-619.

Matt Betts



I study the ways that forest landscape composition and pattern influence animal behavior, species distributions and ecosystem function. As humans are one of the primary drivers of landscape characteristics globally, much of my work is applied and focused on management and conservation, particularly the question of ways of optimizing potential trade-offs between biodiversity and human forest uses. However, understanding mechanisms is key to generalization, so a central part of my research program is basic in nature and links landscape ecology to behavioral ecology, physiology, and evolution. The complexity of ecological systems requires powerful tools that extend beyond ecology into other disciplines - particularly mathematics and computer science. Members of my lab engage actively in such interdisciplinary collaborations.

Selected publications

Benjamin T. Phalan, Joseph M. Northrup, Zhiqiang Yang, Robert L. Deal, Josée S. Rousseau, Thomas A. Spies, Matthew G. Betts (2019) Impacts of the Northwest Forest Plan on forest composition and bird populations. Proceedings of the National Academy of Sciences. DOI:10.1073/pnas.1813072116

Betts, M.G., Wolf, C., Ripple, W.J., Phalan, B., Millers, K.A., Duarte, A., Butchart, S.H.M. and Levi, T. (2017) Global forest loss disproportionately erodes biodiversity in intact landscapes. Nature advance online publication. doi:10.1038/nature23285

Frey, S.J.K., Hadley, A.S., Johnson, S. L., Schulze, M., Jones, J. A. and Betts, M. G. (2016) Spatial models reveal the microclimatic buffering capacity of old-growth forestsScience Advances 2: e1501392.

Kormann, U., Scherber, C., Tscharntke, T., Klein, N., Larbig, Valente, J.M., Hadley, A.S. and Betts, M.G. (2016) Corridors restore animal-mediated pollination in fragmented tropical forest landscapesProceedings of the Royal Society Series B. 283: 1823.

Root, H. and Betts M.G. (2016) Managing moist temperate forests for bioenergy and biodiversityJournal of Forestry 114: 66-74.

Betts, M.G., Hadley, A.S. and Kress, W.J. (2015) Pollinator recognition by a keystone tropical plantProceedings of the National Academy of Sciences of the USA 112: 3433–3438.

Betts, M.G., Fahrig, L., Hadley, A.S., Halstead, K.E., Robinson, W.D. Bowman, J., Wiens, J.A., and Lindenmayer, D.B. (2014) A species-centered approach for uncovering generalities in organism responses to habitat loss and fragmentationEcography 37: 517-527.

Betts, M.G., Verschuyl, J., Giovanini, J., Stokely, T. and Kroll, A.J. (2013) Initial experimental effects of intensive forest management on avian abundanceForest Ecology and Management 310: 1036-1044.

Betts, M.G., Forbes, G.J. and Diamond, A.W. (2007) Thresholds in songbird occurrence in relation to landscape structureConservation Biology 21: 1046-1058.

Kelly Biedenweg

Department of Fisheries and Wildlife


I am an interdisciplinary social scientist who studies how individuals and institutions interact with the natural environment. I explore this primarily from the questions of “How do natural resources contribute to the human experience?” and “How do social relationships influence the management of natural resources?” I spent the first 8 years of my research career in Latin American forests, studying community forestry in the Bolivian Amazon and the Mayan forests as well as environmental leadership in communities in and around Honduran forests. More recently, my research has been focused on the forests and watersheds of the Pacific Northwest. I use tools such as values mapping, cognitive mapping, and the identification and monitoring of cultural ecosystem services.

kelly the birdwatcher.jpg

Selected publications

Biedenweg, K. and N. Gross-Camp (2018) A brave new world: Integrating well-being and conservation. Ecology and Society 23(2):32.

Biedenweg, K., H. Harguth and K. Stiles (2017) The science and politics of human wellbeing: A case study of co-creating indicators for Puget Sound restoration. Ecology and Society 22: 11.

Williams, K., K. Biedenweg and L. Cerveny (2017) Managing for Ecosystem Service Preferences across Residential Classifications near Mt. Baker Snoqualmie National Forest. Forests 8: 157.

Biedenweg, K., T. Scott and R. Scott. (2017) How does engaging with nature relate to life satisfaction? Demonstrating the link between environment-specific social experiences and life satisfaction. Journal of Environmental Psychology 50: 112-124. 

Spalding, A. and K. Biedenweg (2017) Socializing the coast: Engaging the social science of tropical coastal research.  Estuarine, Coastal and Shelf Science 187(5):1-8.

Spalding, A., K. Biedenweg, M. Nelson and A. Hettinger (2017) Demystifying the Human Dimension of Ecological Research. Frontiers in Ecology and the Environment 15(3):119.

Biedenweg, K., K. Stiles and K. Wellman (2016) A holistic framework for identifying human wellbeing indicators for marine policy. Marine Policy 64:31-37.

Besser, D., R. McLain, L. Cerveny, K. Biedenweg and D. Banis (2014) Mapping Landscape Values: Issues, Challenges and Lessons Learned from Field Work on the Olympic Peninsula, Washington. Environmental Practice 16: 138-150. doi: https://doi.org/10.1017/S1466046614000052

Biedenweg, K., L. Cerveny and R. McLain (2014) Values Mapping with Latino Forest Users: Contributing to the dialogue on multiple land use conflict management. Practicing Anthropology 36: 33-37.

Biedenweg, K. and M.C. Monroe (2013)  Teasing Apart the Details: What characteristics of capacity building projects influence community forest management in the Bolivian Amazon?  Human Ecology 41(2): 239-253.

McLain, R., M. Poe, K. Biedenweg, L. Cerveny, D. Besser, and D. Blahna (2013) Making Sense of Human Ecology Mapping: An Overview of Approaches to Integrating Socio-Spatial Data into Environmental Planning. Human Ecology 41(5): 651-665. DOI 10.1007/s10745-013-9573-0

Biedenweg, K. and M.C. Monroe (2013) Cognitive Methods and a Case Study for Assessing Shared Perspectives as a Result of Social Learning. Society and Natural Resources 26(8):931-944. 

Biedenweg, K. and M.C. Monroe (2013) Teasing apart the details: How social learning can affect collective action in the Bolivian Amazon. Human Ecology 41: 239–253.

McLain, R., M. Poe, K. Biedenweg, L. Cerveny, D. Besser and D. Blahna (2013) Making sense of Human Ecology Mapping: An overview of approaches to integrating socio-spatial data into environmental planning. Human Ecology 41: 651–665.

Biedenweg, K. and M.C. Monroe (2013) Cognitive methods and a case study for assessing shared perspectives as a result of social learning. Society and Natural Resources 26: 931-944.

Biedenweg, K. (2012)  A Framework for Evaluating Forest Conservation Implications of Community-based Capacity Building: Experiences from the Northern Bolivian Amazon. Conservation and Society.  

 Barsimontov, J., A. Racelis, K. Biedenweg and M. Digiano (2010)  When Collective Action and Tenure Allocation Collide: Outcomes from community forests in Quintana Roo, Mexico and Petén, Guatemala. Land Use Policy 28(1): 343-352. 

Duchelle, A., K. Biedenweg, C. Lucas, J. Radachowsky, D. Wojcik, M. Londres, D. Alvira, W.L. Bartels, A. Virapongse, K. Kainer (2009) Graduate Students and Knowledge Exchange with Local Stakeholders: Possibilities and Preparation.  Biotropica Special Issue 41:5, 578-587. 

Cabrera, L., K. Biedenweg, I. Camacho, D. Rojas Céspedes, A. Shenkin, D. Elliott, and S. Perz (2009) Las comunidades campesinas y la pavimentación de carreteras en Pando, Bolivia.  Bolivian Studies Journal/RevistaE 8:1

Cerveny, L.C., K. Biedenweg, K. and R. McLain (early online) Mapping meaningful places on Washington's Olympic Peninsula: Toward a deeper understanding of landscape values. Environmental Management.

Badege Bishaw



I am Program Director and Senior Instructor of the Master of Natural Resources and Sustainable Natural Resource Graduate programs at OSU. I have taught courses and developed curricula in International Forestry, Agroforestry and Natural Resources at universities in the United States, South Africa and Ethiopia. As a member of the Sustainable Forest Partnership, I participated in the development of the Sustainable Natural Resources Graduate Certificate at Oregon State University, and more recently, I developed the Forests and Climate Change Graduate Certificate which is taught online through OSU Ecampus. I have over 25 years of experience in forestry, natural resources and agriculture education, research, outreach and administration. I received my PhD in 1993 from Oregon State University in Forest Resources, my MS in 1985 from the University of Dresden, Germany in Tropical Forestry, and my BSc in 1979 from Addis Ababa University, Ethiopia in Plant Sciences.

Selected publications

Bishaw, B., H. Neufeldt, J. Mowo, A. Abdelkadir, J. Muriuki, G. Dalle, T. Assefa, K. Guillozet, H. Kassa, I. K. Dawson, E. Luedeling and C. Mbow (2013) Farmers’ strategies for adapting to and mitigating climate variability and change through agroforestry in Ethiopia and Kenya, edited by C. M. Davis, B. Bernart and A. Dmitriev. Forestry Communications Group, Oregon State University, Corvallis, Oregon.

John Campbell


My research interests center on forest disturbance and its effect on primary production, trophic-transfer, decomposition, and combustion with a special focus on how these processes operate in early-seral communities. Most of my research is conducted in the Pacific coast of North America where harvest, and wildfire, are the predominant agents of disturbance, however, my collaborators and I are also investigating ecosystem function in forests impacted by wind-storms, insect mortality, volcanic eruption, and climate change in various locations throughout North America and the globe. My work is primarily hypothesis-driven involving a combination of field observations and simulation modeling. Whether it be framing questions, interpreting data, or crafting narratives, I am guided equally by a desire to advance theory and inform decision makers. Ride hard, die free.

Selected publications

Campbell, J.L., D.C. Donato, and J.B. Fontaine. 2016. Effects of post-fire logging on fuel dynamics in a mixed-conifer forest, Oregon, USA: a 10-year assessment. International Journal of Wildland Fire. 25 (6), 646-656. PDF

Campbell, J.L. , J.B. Fontaine, and D.C. Donato. 2016. Carbon emissions from decomposition of fire‐killed trees following a large wildfire in Oregon, United States. Journal of Geophysical Research: Biogeosciences 121 (3), 718-730. PDF

Donato, D.C., J.L. Campbell, J.B. Fontaine. 2016 Burning the legacy? Influence of wildfire reburn on dead wood dynamics in a temperate conifer forest. Ecosphere, 7(5). PDF

Meigs, G.W., H.S.J. Zald, J.L. Campbell, W.S. Keeton, and R.E. Kennedy. 2016. Do insect outbreaks reduce the severity of subsequent forest fires? Environmental Research Letters 11 (4), 045008. PDF

Meigs, G.W., J.L. Campbell, H.S.J. Zald, R.E. Kennedy, J.D. Bailey, and D.C. Shaw. 2015. Does wildfire likelihood increase following insect outbreaks in conifer forests? Ecosphere 6(7):118. PDF

Swanson, M.E., N.M. Studevant, J.L. Campbell, and D.C. Donato. 2014. Biological associates of early-seral pre-forest in the Pacific Northwest. Forest Ecology and Management. 324:160-171. PDF

Campbell, J.L., and D.C. Donato. 2014. Trait-based approaches to linking vegetation and food webs in early-seral forests of the Pacific Northwest. Forest Ecology and Management. 324:172-178. PDF

Heckman, K., J.L. Campbell, H. Powers, B.E. Law, and C. Swanston. 2013. The influence of fire on the radiocarbon signature and character of soil organic matter in the Siskiyou National Forest, Oregon, USA. Fire Ecology. 9(2):40-56. PDF

Campbell, J.L., A. Ager. 2013. Forest wildfire, fuel reduction treatments, and Landscape carbon stocks: A sensitivity analysis, Journal of Environmental Management. 121:124-132. PDF

Campbell, J.L., R. Kennedy, W.B. Cohen, and R. Miller. 2012. Regional carbon consequences of Western Juniper encroachment in Oregon. Journal of Rangeland Ecology and Management. 65(3):223-231. PDF

Campbell, J.L., M.E. Harmon, and S.R. Mitchell. 2012. Can fuel reduction treatments really increase forest carbon sequestration by reducing future fire emissions? Frontiers in Ecology and the Environment. 10(2):83-90. PDF

Donato, D.C., J.L., Campbell and J.F. Franklin. 2012. Multiple successional pathways and precocity in forest development: Can some forests be born complex? Journal of Vegetation Science. 23:576-584. PDF

Barger, N.N., A.R. Archer, J.L. Campbell, C. Huang, J.A. Morton, and A.K. Knapp. 2011. Woody plant proliferation in North American drylands: A synthesis of impacts on ecosystem carbon balance. Journal of Geophysical Research. 116, G00K07, doi:10.1029/2010JG001506. PDF

Meigs, G.W. and J.L. Campbell. 2010. Comment on “Prescribed fire as a means of reducing forest carbon emissions in the Western United States”. Environmental Science and Technology. 44: 6520. PDF

Halofsky, J. D. Donato, D. Hibbs, J. Campbell, M. Cannon, J. Fontaine, J. Thompson, R. Anthony, L. Kayes, B. Law, D. Peterson, and T. Spies. 2010. Mixed-Severity Fire Regimes: Lessons from the Klamath-Siskiyou Ecoregion. Ecosphere, 2(4): Article 40. PDF

Fontaine, J.B., D.C. Donato, J.L. Campbell, J.G. Martin, and B.E. Law. 2010. Effects of post-fire logging on forest surface air temperatures in the Siskiyou Mountains, Oregon, USA. Forestry. doi:10.1093/forestry/cpq030. PDF

Duane, M., W.B. Cohen, J.L. Campbell, and T. Hudiburg. 2009. Implications of two different field sampling designs on Landsat-based maps of forest attributes. Forest Science. 56(4): 405-416. PDF

Donato, D.C., J.L. Campbell, J.B. Fontaine, B.E. Law. 2009. Quantifying char in post-fire woody detritus inventories. Journal of Fire Ecology. 5(2):104-115. PDF

Hudiburg, T.M., B.E. Law, D.P. Turner, J. Campbell, D. Donato, and M. Duane. 2009. Carbon dynamics of Oregon and Northern California forests and potential land-based carbon storage. Ecological Applications. 19(1): 163-180. PDF

Donato, D.C., J.B. Fontaine, J.L. Campbell, W.D. Robinson, J.B. Kauffan, and B.E. Law. 2009. Conifer regeneration in stand-replacement portions of a landscape-scale mixed-severity wildfire. Canadian Journal of Forest Research. 39: 823-838. PDF

Meigs, G.W., B.E. Law, D.C. Donato, J.L. Campbell, and J. Martin. 2009. Influence of mixed-severity wildfires on pyrogenic carbon transfers, postfire carbon balance, and regeneration trajectories in the eastern Cascades, Oregon. Ecosystems. 12:1246-1267. PDF

Campbell, J.C., G. Alberti, and B.E. Law. 2008. Carbon dynamics of a Ponderosa pine plantation following fuel reduction treatment in the northern Sierra Nevada. Forest Ecology and Management. 275:453-463. PDF

Campbell, J.C., D.C. Donato, D.A. Azuma, and B.E. Law. 2007. Pyrogenic carbon emission from a large wildfire in Oregon, USA. Journal of Geophysical Research-Atmospheres. 112(G4): No. G04014. PDF

Turner, P. D. Ritts, E. Law, B. Cohen, Z. Yang, T. Hudiburg, L. Campbell, and M. Duane. 2007. Scaling net ecosystem production and net biome production over a heterogeneous region in the western United States. Biogeosciences. 4(4): 597-612. PDF

Donato, D.C., J.B. Fontaine, J.L. Campbell, W.D. Robinson, J.B. Kauffman, and B.E. Law. 2006. Response to Comments on “Post-wildfire logging hinders regeneration and increased fire risk”. Science. 313: 615c. PDF

Donato, D.C., J.B. Fontaine, J.L. Campbell, W.D. Robinson, J.B. Kauffman, and B.E. Law. 2006. Post-wildfire logging hinders regeneration and increased fire risk. Science. 311(4759): 352. PDF

Loescher, H, B.E. Law, L. Mahrt, D.Y. Hollinger, J.L. Campbell, and S.C. Wofsy. 2006. Uncertainties in, and interpretation of, carbon flux estimates using the eddy covariance technique. Journal of Geophysical Research-Atmospheres. 111, D21S90, doi:10.1029/2005JD006932. PDF

Law, B.E., D. Turner, J.L. Campbell, M. Lefsky, M. Guzy, O. Sun, S. VanTuyl, and W.B. Cohen. 2006. Carbon fluxes across regions: Observational constraints at mutable scales. In: J. Wu, K.B. Jones, H. Li, and O.L. Loucks (eds.) Scaling and Uncertainty Analysis in Ecology: Methods and Applications. Pp 163-186. Springer, Netherlands. PDF

Campbell, J.L. and B.E. Law. 2005. Forest soil respiration across three climatically-distinct chronosequences in Oregon. Biogeochemistry. 73(1):109-125. PDF

Turner, D.P., W. Ritts, W.B. Cohen, et al. 2005. Site-level evaluation of satellite-based global NPP/GPP monitoring. Global Change Biology. 11:1-19. PDF

Campbell, J.L., O.J. Sun and B.E. Law. 2004. Disturbance and net ecosystem production across three climatically distinct landscapes. Global Biogeochemical Cycles. 18:GB4017. PDF

Campbell, J.L., O.J. Sun and B.E. Law. 2004. Supply-side controls on soil respiration among Oregon forests. Global Change Biology. 10:1857-1869. PDF

Law, B.E., D. Turner, J.L. Campbell, O.J. Sun, S. VanTuyl, W.D. Ritts and W.B. Cohen. 2004. Disturbance and Climate Effects on Carbon Stocks and Fluxes Across Western Oregon USA. Global Change Biology. 10:1429-1444. PDF

Sun, O.J., J.L. Campbell and B.E. Law. 2004. Dynamics of carbon storage in soils and detritus across chronosequences of different forest types in the Pacific Northwest, USA. Global Change Biology. 10:1470-1481. PDF

Law, B.E., O.J. Sun, J.L. Campbell, S. VanTuyl and P. Thornton P. 2003. Changes in carbon storage and fluxes in a chronosequence of ponderosa pine. Global Change Biology, 9, 510-524. PDF

Gower S.T., A. Hunter, J.L. Campbell, J. Vogel, H. Veldhuis, J. Harden, S. Trumbore, J.M. Norman and C.J. Kucharik. 2000. Nutrient dynamics of the southern and northern BOREAS boreal forests. Ecoscience 7 (4):481-490. PDF

Campbell, J.L. and S.T. Gower. 2000. Detritus production and soil N transformations in old-growth eastern hemlock and sugar maple stands. Ecosystems 3(2): 185-192. PDF

Jason Dunham

DEPT. of fisheries and wildlife

Supervisory Research Ecologist
Forest and Rangeland Ecosystem Science Center
U.S. Geological Survey

Jason_Elks_Nov2017 (2) (1).JPG

My research is focused on landscape ecology of aquatic ecosystems, conservation biology of focal species, ecology of natural disturbance, biological invasions, and monitoring.  My current research is focused on climate adaptation, local and regional effects of forest and rangeland management on streams, conservation planning for wetlands in arid ecosystems, ecology of beaver and beaver-assisted restoration, and human dimensions of riverscapes.

Selected publications

JB Dunham, BE Rieman. 1999. Metapopulation structure of bull trout: influences of physical, biotic, and geometrical landscape characteristics. Ecological Applications 9:642-655.

SJ Wenger, DJ Isaak, CH Luce, HM Neville, KD Fausch, JB Dunham, et al. 2011. Flow regime, temperature, and biotic interactions drive differential declines of trout species under climate change. Proceedings of the National Academy of Sciences 108:14175-14180.

BE Rieman, JB Dunham. 2000. Metapopulations and salmonids: a synthesis of life history patterns and empirical observations. Ecology of Freshwater Fish 9:51-64.

JB Dunham, SB Adams, RE Schroeter, DC Novinger. 2002. Alien invasions in aquatic ecosystems: toward an understanding of brook trout invasions and potential impacts on inland cutthroat trout in western North America. Reviews in Fish Biology and Fisheries 12:373-391.

HM Neville, JB Dunham, MM Peacock. 2006. Landscape attributes and life history variability shape genetic structure of trout populations in a stream network. Landscape Ecology 21:901-916.

Clinton Epps

associate professor
DEPT. of fisheries and wildlife


Research in my lab includes the study of dispersal and connectivity, fragmented populations, metapopulation dynamics, effects of climate change, wildlife disease, wildlife management, and the application of population genetics to estimating dispersal and other population parameters.

Selected publications

Castillo, J. A., C. W. Epps, A. R. Davis, & S. A. Cushman. 2014. Landscape effects on gene flow for a climate-sensitive montane species, the American pika. Molecular Ecology 23:843-856.

Epps, C. W., J. D. Wehausen, W. B. Sloan, S. Holt, T. G. Creech, R. S. Crowhurst, J. R. Jaeger, K. M. Longshore, & R. J. Monello. In press. Fifty years after Welles and Welles: distribution and genetic structure of desert bighorn sheep in Death Valley National Park. Proceedings of the 1st Annual Death Valley Natural History Conference, Death Valley National Park, California.

Epps, C. W., S. K. Wasser, J. L. Keim, B. M. Mutayoba, & J. S. Brashares. 2013. Quantifying past and present connectivity illuminates a rapidly changing landscape for the African elephant. Molecular Ecology 22:1574-1588.

Epps, C. W., J. A. Castillo, A. Schmidt-Küntzel, P. du Preez, G. Stuart-Hill, M. Jago, & R. Naidoo. 2013. Contrasting historical and recent gene flow among African buffalo herds in the Caprivi Strip of Namibia. Journal of Heredity 104:172-181.

Epps, C.W., B. M. Mutayoba, L. E. Gwin, & J. S. Brashares. 2011. An empirical evaluation of the African elephant as a focal species for connectivity planning East Africa. Diversity and Distributions 17:603-612. 

Sarah Frey



My interest is in quantitative population ecology, with a focus on understanding how environmental patterns at different scales affect biodiversity in both temperate and tropical forest systems. Specifically, I study how land-use and climate changes in montane forest landscapes influence species distributions, abundance trends, and community composition. I apply novel analytical tools (i.e., hierarchical models, machine learning) to address my research questions. Some of my recent research has examined 1) the potential for vegetation structure to provide microrefugia for biodiversity in the face of regional warming in the Pacific Northwest, 2) the influence of tropical forest fragmentation on hummingbird community structure and functional traits in Costa Rica, and 3) how scale and imperfect detection influence long term abundance trends of forest birds at Hubbard Brook.

Selected publications

Complete list of publications

Hadley, A.S., S.J.K. Frey, W.D. Robinson, and M.G. Betts (2017) Forest fragmentation and loss reduce richness, availability and specialization in tropical hummingbird communities. Biotropica: doi: 10.1111/btp.12487

Frey, S.J.K., A.S. Hadley, S.L. Johnson, M. Schulze, J. Jones, and M.G. Betts (2016) Spatial models reveal microclimatic buffering capacity of old-growth forest. Science Advances 2: e1501392.

Frey, S.J.K., A.S. Hadley, and M.G. Betts (2016) Microclimate predicts within-season distribution dynamics of montane forest birds. Diversity and Distributions 22: 944-959.

Tiffany Garcia

DEPT. Of Fisheries and wildlife


My research encompasses a wide variety of biological disciplines, including aquatic, behavioral and community ecology, invasion biology, conservation biology, and herpetology.  I typically work in freshwater habitats within forested landscapes and address ecologically based questions using amphibian and invertebrate systems.  I have projects quantifying predator/prey dynamics, competition regimes, stressor defense strategies, and microhabitat preference using observational, theoretical and empirical methods.  I enjoy incorporating both laboratory and field components in my work, and appreciate a diverse working environment.  

Selected publications

Kroll, A.J., T.S. Garcia, J. Jones, K. Dugger, B. Murden, J. Johnson, S. Peterman, M. Brintz and M. Rochelle (2015) Evaluating multi-level models to assess occupancy state responses of Plethodontid salamandersPLoS ONE 10: e0145899.

Garcia, T.S., J.C. Rowe and J.B. Doyle (2015) A tad too high: Sensitivity to UV-B radiation may limit invasion potential of America bullfrogs in the Pacific Northwest invasion rangeAquatic Invasions 10: 237-247.

Thurman, L.L., T.S. Garcia and P. Hoffman (2014) Elevational differences in trait response to UV-B radiation by long-toed salamander populationsOecologia 175: 835-845.

Bucciarelli, G.M., A.R. Blaustein, T.S. Garcia, and L.B. Kats (2014) Invasion Complexities: The diverse impacts of invasive species on amphibiansCopeia 2014: 611-632.

Rowe, J.C. and T.S. Garcia (2013) Impacts of wetland restoration efforts on an amphibian assemblage in a multi-invader communityWetlands 34: 141-153.

Baker, N.J., B. Bancroft, and T.S. Garcia (2013) A meta-analysis of the effects of pesticides and fertilizers on amphibiansScience of the Total Environment 449: 150-156.

Cook, M.T., S.S Heppell, and T.S. Garcia (2013) Invasive bullfrog larvae lack developmental plasticity to changing hydroperiodJournal of Wildlife Management 77: 655-662.

Garcia, T.S., L.L. Thurman, J.C. Rowe, and S. Selego (2012) Antipredator behavior of American bullfrogs (Lithobates catesbeianus) in a novel environment. Ethology 118: 1-9.

Rowe, J.C., and T.S. Garcia (2012) First record of predation by the invasive American bullfrog (Lithobates catesbeinus) on the threatened Oregon chub (Oregonichthys crameri). Herpetological Review 43: 633-634.

Hanshew, B.A. and T.S. Garcia (2012) Invasion of the shelter snatchers: shelter use dynamics of co-occurring native and invasive crayfish from Oregon's Willamette ValleyFreshwater Biology 57: 2285-2296.

Adam Hadley



I am especially interested in the effects of landscape disturbances on ecological processes. My research efforts focus on the intersection of landscape ecology, pollination ecology and behavioral ecology. My current research is being conducted in two study systems, one located in tropical premontane forest of Southern Costa Rica and the other in a temperate system at the H. J. Andrews Experimental Forest in Oregon. My general interests include: 1) Animal movements and how they are influenced by landscape disturbances. 2) Landscape effects on plant and animal interactions (e.g. pollination, seed dispersal, stability of ecological networks). 3) Landscape genetics. 4) Drivers of species distributions. And 5) The role of social information in resource and habitat selection.

Selected publications

Hadley, A. S. and M. G. Betts. (2016) Refocusing habitat fragmentation research using lessons from the last decade. Current Landscape Ecology Reports 1: 55-66.

Frey, S. J. K., A. S. Hadley, S. L. Johnson, M. Schulze, J. Jones, and M. G. Betts (2016) Spatial models reveal microclimatic buffering capacity of old-growth forest. Science Advances 2: e1501392.

Betts, M. G., A. S. Hadley, and W. J. Kress. (2015) Pollinator recognition by a keystone tropical plant. Proceedings of the National Academy of Sciences of the United States of America 112: 3433-3438.

Hadley, A. S., S. J. K. Frey, W. D. Robinson, W. J. Kress, and M. G. Betts. (2014) Tropical forest fragmentation limits pollination of a keystone understory herb. Ecology 95: 2202-2212.

Hadley, A. S., and Betts, M. G. (2012) The effects of landscape fragmentation on pollination dynamics: absence of evidence not evidence of absence. Biological Reviews 87: 526-544.

Joan C. Hagar

USGS Forest & Rangeland Ecosystem Science Center


I conduct research on wildlife habitat relationships in forests, woodlands, and rangelands, including investigations of the effects of natural disturbance and management strategies on wildlife habitat, and evaluation of the effectiveness of habitat restoration treatments for wildlife.

Selected publications

All available at: https://fresc.usgs.gov/people/Profile.aspx?Emp_ID=34

Hagar, J.C. 1999. Influence of riparian buffer width on bird assemblages in western Oregon. J. Wildlife Management 63(2): 484-496.

Hagar, J.C., and Stern, M.A., 2001. Avifauna in oak woodlands of the Willamette Valley, Oregon. Northwestern Naturalist 82: 12 – 25.

Hagar, J.C., L.M. Ganio, and S. Howlin. 2004. Short-term response of songbirds to experimental thinning of young Douglas-fir forests in the Oregon Cascades. Forest Ecology and Management 199: 333-347.

Hagar, J.C., 2007. Wildlife species associated with non-coniferous vegetation in Pacific Northwest conifer forests- A review. Forest Ecology and Management 246: 108-122.

Hagar, J.C., 2007, Key elements of stand structure for wildlife in production forests west of the Cascade Mountains. In Harrington, T.B., Nicholas, G.E., eds., Managing for Wildlife Habitat in Westside Production Forests: PNW-GTR-695, Portland, OR, U.S. Forest Service Pacific Northwest Research Station, pp. 35-48


Reem Hajjar



As a social scientist with training in ecology, I study the relationship between forests and livelihoods, and how various governance mechanisms (policies, norms, and markets) shape that relationship. Most of my work to-date has taken place in tropical and sub-tropical forests, with a recent extension into the Pacific Northwest of North America. A driving factor behind my research is figuring out how governance and decision-making processes around forests can be made more just, participatory, and driven from the bottom-up, so that they effectively reflect realities on the ground, and the realities of often marginalized forest-dependent people. I aim to answer questions such as, can forests be sustainably used as pathways to prosperity? If yes, by which mechanisms? What are the livelihood and landscape impacts of tenure and forest policy changes, and how can these changes result in more equitable outcomes? How transferable are governance mechanisms from one context to another? Most recently, I’ve been working on projects related to community-based forest management in Mexico, global reviews on community forestry and small-scale forest enterprises, gendered impacts of land grabs in Ethiopia, impacts of forest policy changes in Ghana, and cattle sector certification in Brazil.

Selected publications

Recent publications: Google scholar

Phalan, B, R Hajjar. Developing without deforestation. Nature Plants – News & Views. 3: 17120.

Bogaerts, M, L Cirighiri, I Robinson, M Rodkin, R Hajjar, C Costa Junior, P Newton. 2017. Climate change mitigation through intensified pasture management: Estimating greenhouse gas emissions on cattle farms in the Brazilian Amazon. Journal of Cleaner Production 162: 1539-1550.

Tomaselli, MF, R Hajjar, AE Ramon Hidalgo, A Vasquez. 2017. The problematic old roots of new green narratives: How far can they take us in re-imagining sustainability in the forestry sector? Special Issue on the Green Economy in International Forestry Review 19 (S1): 1-13.

Hajjar, R, RA Kozak. 2017. The evolution of forest producer associations and their current role in REDD+: Case studies from Quintana Roo, Mexico. Land Use Policy 60: 373-383

Hajjar, R, JO Oldekop, P Cronkleton, E Etue, P Newton, AJ Russell, JS Tjajadi, W Zhou, A Agrawal. 2016. The data not collected on community forestry. Conservation Biology 30: 1357-1262.

Hajjar, R, O Sanchez-Badini, RA Kozak. 2016. Promoting small and medium forest enterprises in national REDD+ strategies: A multi-country analysis of enabling environments. Climate Policy 17: 731-763.

Hajjar, R, RA Kozak. 2015. Exploring public perceptions of forest adaptation strategies in Western Canada: Implications for Policy-makers. Forest Policy and Economics 61: 59-69.

Hajjar, R. 2015. Advancing small-scale forestry under FLEGT and REDD in Ghana. Forest Policy and Economics 58: 12-20.

Rebecca Hutchinson

School of Electrical Engineering & Computer Science
Department of Fisheries & Wildlife


My research is at the intersection of machine learning and ecology. I am part of the computational sustainability community, trying to find ways that computer science can contribute to promoting the health of the Earth's ecosystems and bringing interesting new problems back to computer science. Much of my work is on computational methods for species distribution modeling. I work primarily with hierarchical latent variable models that represent both ecological and observation processes; for example, occupancy models and their variants fall within this paradigm. My current research is on robust parameter estimation methods for these models and techniques for incorporating semi-parametric techniques into probabilistic models. I am also interested in methods for analyzing species interaction networks and strategies for evaluating species distribution models.

Selected publications

Valente, J.J., Hutchinson, R.A. and Betts, M.G., (online early) Distinguishing distribution dynamics from temporary emigration using dynamic occupancy models. Methods in Ecology and Evolution doi:10.1111/2041-210X.12840

Prudic, K.L., McFarland, K.P., Oliver, J.C., Hutchinson, R.A., Long, E.C., Kerr, J.T. and Larrivée, M. (2017) eButterfly: Leveraging Massive Online Citizen Science for Butterfly Conservation. Insects 8: 53.

Hutchinson, R.A., Valente, J.J., Emerson, S.C., Betts, M.G. and Dietterich, T.G. (2015) Penalized likelihood methods improve parameter estimates in occupancy models. Methods in Ecology and Evolution 6: 949–959. doi:10.1111/2041-210X.12368

Shirley, S.M., Yang, Z., Hutchinson, R.A., Alexander, J.D., McGarigal, K. and Betts, M.G. (2013) Species distribution modelling for the people: unclassified landsat TM imagery predicts bird occurrence at fine resolutions. Diversity and Distributions 19: 855–866. doi:10.1111/ddi.12093

Hochachka, W.M., Fink, D., Hutchinson, R.A., Sheldon, D., Wong, W.-K. and Kelling, S. (2012) Data-intensive science applied to broad-scale citizen science. Trends in Ecology & Evolution 27: 130–137. doi:10.1016/j.tree.2011.11.006

Andy Jones

DEPT. OF Botany and Plant Pathology


The Jones lab is focused on understanding the mechanisms responsible for the origin and maintenance of plant species and genetic diversity.  Projects employ a variety of field, lab, and bioinformatic approaches across population, community, and landscape scales.  We are particularly interested in developing and applying novel molecular ecological approaches towards understanding plant population and community dynamics in the recent and ancient past to environmental change and it's implications for ecosystem responses in the future.  Current projects include understanding the extent to which tropical tree species may be locally adapted to rainfall regimes across the Isthmus of Panama; the effect of the 2015 El Nino associated drought on tropical tree seedling regeneration in Panama; how habitat fragmentation may alter hummingbird-mediated pollen movement and gene flow in Heliconia tortuosa in Costa Rica; developing genomic resources for threatened tropical timber species Cedrela odorata in Peru; and exploring the role of plant pathogens in the origin and maintenance of tree diversity in a Pacific Northwest old growth forest.

Selected publications

Barberán, A., K.L. McGuire, J.A. Wolf, F.A. Jones, S.J. Wright, B.L. Turner, A. Essene, S.P. Hubbell, B. Faircloth and N. Fierer (2015) Relating belowground microbial composition to the taxonomic, phylogenetic, and functional trait distributions of trees in a tropical forestEcology Letters 18: 1397-1405.

Jones, F.A., I. Ceron-Souza, B.D. Hardesty, and C.W. Dick (2013) Genetic evidence of Quaternary demographic changes in four rainforest tree species sampled across the Isthmus of Panama. Journal of Biogeography 40: 720-731.

Jones, F.A., D.L. Erickson, M.A. Bernal, W.J. Kress, E.A. Herre, H. Muller-Landau, O. Sanjur, E. Bermingham, and B.L. Turner (2011) The roots of diversity: below ground species overlap in a tropical forest revealed by DNA barcoding and inverse modeling of root distributions. PLoS ONE 6: e24506.

Jones, F.A. and L.S. Comita (2008) Neighbourhood density and genetic relatedness interact to determine fruit set and abortion rates in a continuous tropical tree population. Proceedings of the Royal Society of London Series B 275: 2759-2767. 

Jones, F.A., J. Chen, G.-J. Weng, and S.P. Hubbell (2005) A genetic evaluation of seed dispersal in the Neotropical tree, Jacaranda copaia (Bignoniaceae). The American Naturalist 166: 843-855.

Julia Jones

dept. of Geography


Specialty: Spatio-temporal statistics, forest hydrology, landscape ecology, surface processes, biogeography.

Research Interests: Land use, climate change and disturbance effects on hydrology, geomorphology, water quality


Selected publications

Landscape patterns and diversity of meadow plants and flower-visitors in a mountain landscape. J.A. Jones, R. Hutchinson, A. Moldenke, V. Pfeiffer, E. Helderop, E. Thomas, J. Griffin, A. Reinholtz. Landscape Ecology. (2018)

Forest harvest legacies control dissolved organic carbon export in small watersheds, western Oregon. K. Lajtha, J.A. Jones. Biogeochemistry,140, 299-315. (2018)

Forest landscape hydrology in a ‘new normal’ era of climate and land use change. Jones JA and Wei X, Coordinating lead authors. Pp. 81-100 in Creed IF and van Noordwijk M (eds.). Forests and Water on a Changing Planet: Vulnerability, Adaptation and Governance Opportunities. A Global Assessment Report. IUFRO World Series Volume 38. Vienna. 192 p. (2018)

Climate-Forest-Water-People Relations: Seven System Delineations. van Noordwijk M, Creed IF, Jones JA and Wei Xi, Coordinating lead authors. Pp. 27-60 in Creed IF and van Noordwijk M (eds.), 2018. Forests and Water on a Changing Planet: Vulnerability, Adaptation and Governance Opportunities. A Global Assessment Report. IUFRO World Series Volume 38. Vienna. 192 p. (2018)

Forests and water in South America, Jones, J., Almeida, A., Cisneros, F., Iroumé, A., Jobbágy, E., Lara, A., Lima, W. d. P., Little, C., Llerena, C., Silveira, L., and Villegas, J.C. Hydrological Processes, doi: 10.1002/hyp.11035 (2017)

Urs G. Kormann



I am a Postdoc with Matt Betts at Oregon State University. I seek to understand the dynamics of biotic communities, interactions networks and ecosystem functions in our rapidly changing world.  I explore ways to implement this knowledge for biodiversity conservation in human-modified systems. I am particularly interested in synergistic effects of global change drivers (e.g. habitat loss, land use intensification,  climate change) in space and time.  Broadly speaking,  my interests encompass (i) why organisms are where they are, (ii) how and why species interact(or not),  (iii) what ecosystem functions they provide (or not) and (iv) movement ecology. I like to tackle research questions from different angles, that is,  by combining experimental, observational, genetic and statistical modelling approaches. I have been working in tropical and temperate systems on a broad range of taxa, sometimes with large community datasets, sometimes single-species data, but with a certain taxonomic bias towards birds, insects and plants. Currently I am involved in two projects; effects ofintensive forest management on biodiversity, ecosystem functioning and pollinator-plant interaction webs in North America , and landscape effects on plant-animal interactions in Southern Costa Rica. I have an active interest in statistics, particularly in hierarchical modelling and statistical techniques to assess community shifts in space and time.

Selected publications

Kormann, U., Scherber, C., Tscharntke, T., Klein, N., Larbig, M., Valente, JV., Hadley AS and Betts MG. (2016) Corridors restore animal-mediated pollination in fragmented tropical forest landscapes. Proc. R. Soc. B 283: 20152347

Zahawi, R. A., Duran, G., & Kormann, U. (2015) Sixty-seven years of land-use change in southern Costa Rica. PloS ONE 10(11): e0143554.

Pfeifer, M., [...], Kormann, U., et. al.  (2013) BIOFRAG – A new database for analysing BIOdiversity responses to forest FRAGmentation. Ecology and Evolution 4(9): 1524 – 1537.

Kormann, U., Gugerli, F., Ray, N., Excoffier, L., & Bollmann, K. (2012) Parsimony-based pedigree analysis and individual-based landscape genetics suggest topography to restrict dispersal and connectivity in the endangered capercaillie. Biological Conservation 152: 241-252.

Kormann, U., Rösch, V, Batary, P., Tscharntke, T., Orci, KM., Samu, F., & Scherber, C. (2015) Local and landscape management affects trait-mediated biodiversity of nine taxa on small grassland fragments. Diversity and Distributions 21: 1204-1217.

Meg Krawchuk

DEPT. OF forest ecosystems & society


My research and teaching focuses on landscape and fire ecology, pyrogeography, and conservation science. I work at scales from local to global, addressing the drivers and effects of ecological disturbances with a primary interest in wildfire. Recent investigations include: i) structure, function, and location of fire refugia within burn mosaics of western North American forest ecosystems, ii) ecological implications of overlapping short-interval disturbances such as insect outbreaks, forest harvest, and wildfire, iii) spatially varying constraints over modern patterns of burning, iv) theory and tools necessary for science-based conservation planning in boreal forest ecosystems of Canada and the United States. We’ve just begun a project studying the effects of the invasive grass Ventenata dubia on fire regimes and ecology of Oregon’s Blue Mountain Ecoregion. My research group integrates field and computing studies with expertise shared among undergraduate, graduate, and post-graduate researchers. All of my work is anchored through effective collaborations and motivated by the enjoyment of science.  

Selected publications

Influence of fire refugia spatial pattern on post-fire forest recovery in Oregon’s Blue Mountains.William M. Downing, Meg A. Krawchuk, Garrett W. Meigs, Sandra L. Haire, Jonathan D. Coop, Ryan B. Walker, Ellen Whitman, Geneva Chong, Carol Miller. 2019. Landscape Ecology.

Fire refugia: what are they and why do they matter for global change? Meddens, A., Kolden, C., Lutz, J., Smith, A., Cansler, C., Abatzoglou, J., Meigs, G., Downing, W., and Krawchuk, M.A. 2018. BioScience 11.

Composition and structure of forest fire refugia: what are the ecosystem legacies across burned landscapes. Meigs, G., and Krawchuk, M.A. 2018. Forests 2018, 9(5), 243.

Wildfires managed for restoration enhance ecological resilience. Barros, A., Ager, A., Day, M., Krawchuk, M.A., and Spies, T.A. 2018. Ecosphere 9:3 

Adapt to more wildfire in western North American forests as climate changes. Schoennagel, T., Balch, J.K., Brenkert-Smith, H., Dennison, P.E., Harvey, B.J., Krawchuk, M., Mietkiewicz, N., Morgan, P., Moritz, M.A., Rasker, R., Turner, M.G., Whitlock, C. 2017. Proceedings of the National Academy of Sciences 114:4582-4590

Spatially varying constraints of human-caused fire occurrence in British Columbia, Canada. Camp, P., and Krawchuk, M.A. 2017. International Journal of Wildland Fire. 26:219-229

Topographic and fire weather controls of fire refugia in forested ecosystems of northwestern North America. Krawchuk, M.A., Haire, S., Coop, J.C., Parisien, M-A., Whitman, E., Chong, G., Miller, C. 2016. Ecosphere 7(12) e01632

The effects of disturbance history on ground-layer plant community composition in British Columbia. Ton, M., and Krawchuk, M.A. 2016. Forests 7(5) 109

Future changes in climatic water balance determine potential for transformational shifts in Australian fire regimes. Boer, M.M., Bowman, D.M.J.S., Murphy, B.P., Cary, G.J., Cochrane, M.A., Fensham, R.J., Krawchuk, M.A., Price, O.F., Resco De Dios, V., Williams, R.J., Bradstock, R.A. 2016. Environmental Research Letters 11 065002

Burning issues: statistical analyses of global fire to inform assessments of environmental change. Krawchuk, M.A., Moritz, M.A. 2014. Environmetrics 25:472-481.

Climate change and disruption in global fire activity. Moritz, M.A., Parisien, M-A., Batllori, E., Krawchuk, M.A., Van Dorn, J., Ganz, D., and Hayhoe, K. 2012. Ecosphere 3:art49. 

Global protected areas and IUCN designations: does the category match the condition? Leroux, S.J., Krawchuk, M.A., Schmiegelow, F.K.A., Cumming, S.G., Lisgo, K., Anderson, L., and Petkova, M. 2010. Biological Conservation 143:609-616.

Fire in the Earth System. Bowman, D.M.J.S., Balch, J.K., Artaxo, P., Bond, W.J., Carlson, J.M., Cochrane, M.A., D’Antonio, C.M., DeFries, R.S., Doyle, J.C., Harrison, S.P., Johnston, F.H., Keeley, J.E., Krawchuk, M.A., Kull, C.A., Marston, J.B., Moritz, M.A., Prentice, I.C., Roos, C.I., Scott, A.C., Swetnam, T.W., van der Werf, G.R., and Pyne, S.J. 2009. Science 324:481-484.

Application of mixed models to the study of resource selection by animals. Gillies, C., Hebblewhite, M., Nielsen, S., Krawchuk, M.A., Aldridge, C., Frair, J., Saher, J., Stevens, C., and Jerde, C. 2006. Journal of Animal Ecology 75:837-1044.

Beverly Law


College of Earth, Ocean & Atmospheric Sciences


I research how climate and disturbance affect ecosystem function in forests, woodlands, and shrublands at scales from leaf to ecosystem, region and the globe. Human influence on ecosystems is a major driver of change. My work includes the effects of management activities such as thinning and land-use change on ecosystem function, and studies on species-specific traits that affect sensitivity to climate. My research team conducts field measurements along ecoclimatic gradients on drought sensitivity of species and developmental stages of trees, ecosystem-atmosphere exchange of carbon, water and energy in different forest types and age classes and in sagebrush steppe. We combine observations and land-system modeling that incorporates physiological trait sensitivity to predict vulnerability of ecosystems to changing climate in the future, and the interactive effects of climate and actions taken to mitigate impacts in Oregon and the western US. We emphasize the importance of conservation of biodiversity and ecosystem function in the face of climate extremes and increasing human appropriation of land and productivity in the Anthropocene.

Selected publications

Berner, L.T., B.E. Law, A. Meddens and J. Hicke (2017) Tree biomass mortality from fires, bark beetles, and timber harvest during a hot, dry decade in the western United States (2003-2012). Environmental Research Letters 103485.

Berner, L.T. and B.E. Law (2015) Water limitations on forest carbon cycling and conifer traits along a steep climatic gradient in the Cascade Mountains, Oregon. Biogeosciences 12: 6617-6635. doi:10.5194/bg-12-6617-2015.

Law, B.E. and R.H. Waring (2015) Carbon implications of current and future effects of drought, fire and management on Pacific Northwest forests. Forest Ecology and Management 355: 4-14. dx.doi.org/10.1016/j.foreco.2014.11.023.

Law, B.E. (2014) Regional analysis of drought and heat impacts on forests: current and future science directions. Global Change Biology 20: 3595-3599. doi: 10.1111/gcb.12651.

Law, B.E., T. Hudiburg and S. Luyssaert (2013) Thinning effects on forest productivity: Consequences of preserving old forests and mitigating impacts of fire and drought. Plant Ecology & Diversity 6: 73-85. John Grace Festschrift - Forest and Environment Special Issue. doi: 10.1080/17550874.2012.679013.

Gonçalves, F., M. Yatskov, J.R. dos Santos, R.N. Treuhaft and B.E. Law (2010) Relating LANDSAT ETM+ and forest inventory data for mapping successional stages in a tropical wet forest. Ambiência Guarapuava (PR) v.6 Ed. Especial: 167 – 174. ISSN 1808 – 0251.

Fontaine, J.B., Donato, D., W.D. Robinson, B.E. Law and J.B. Kaufmann (2009) Bird communities following high-severity fire: Response to single and repeat fires in a mixed-evergreen forest, Oregon, USA. Forest Ecology and Management 257: 1496-1504.

Donato, D., J.B. Fontaine, W.D. Robinson, J.B. Kaufmann and B.E. Law (2009) Vegetation response to a short interval between high-severity wildfires in a mixed-evergreen forest. Journal of Ecology 97: 142-154. doi: 10.1111/j.1365-2745.2008.01456.x

Turner, D.P., M. Guzy, M. Lefsky, S. Van Tuyl, O. Sun, C. Daly and B.E. Law (2003) Effects of land use and fine scale environmental heterogeneity on net ecosystem production over a temperate coniferous forest landscape. Tellus 55B: 657-668.

Law, B.E., E. Falge, D.D. Baldocchi, P. Bakwin, P. Berbigier, K. Davis, A.J. Dolman, M. Falk, J.D. Fuentes, A. Goldstein, A. Granier, A. Grelle, D. Hollinger, I.A. Janssens, P. Jarvis, N.O. Jensen, G. Katul, Y. Mahli, G. Matteucci, R. Monson, W. Munger, W. Oechel, R. Olson, K. Pilegaard, K.T. Paw U, H. Thorgeirsson, R. Valentini, S. Verma, T. Vesala, K. Wilson and S. Wofsy (2002) Environmental controls over carbon dioxide and water vapor exchange of terrestrial vegetation. Agricultural and Forest Meteorology 113: 97-120. *World Meteorological Organization Paper of the Year, Norbert-Gerbier Mumm Award.

Damon Lesmeister

Pacific Northwest Research Station
USDA Forest Service

DEPT. OF Fisheries and wildlife

My current research focuses on conservation and management problems in biology and ecology of wildlife species associated with late-successional forests in the Pacific Northwest. In my lab we directly or indirectly address questions related to interactions between avian and mammalian predators and their prey. We are particularly interested in long-term studies of demography, effects of disturbance, spatial ecology, resource selection, and linkages between predator and prey populations. Current projects include long-term demography of northern spotted owls; long-term abundance and survival of small mammals in an old-growth forest; experimentally testing hypotheses of factors that limit red tree vole occupancy of young forest; ecology of small carnivores in an old-growth forest; testing autonomous recording units to detect rare forest owls; mixed-severity wildfire effects on northern spotted owl occupancy dynamics, habitat, and prey; and distribution modeling of an imperiled red tree vole population.   

SELECTED Publications

Glenn, E. M., D. B. Lesmeister, R. J. Davis, B. Hollen, A. Poopatanapong. 2017. Estimating density of a territorial species in a dynamic landscape. Landscape Ecology 32(3): 563–579.

Gompper, M. E., D. B. Lesmeister, J. C. Ray, J. R. Malcolm, and R. W. Kays. 2016. Differential habitat use or intraguild interactions: What structures a carnivore community? PLoS ONE 11: e0146055.

Lesmeister, D. B., C. K. Nielsen, E. M. Schauber, and E. C. Hellgren. 2015. Spatial and temporal structure of a mesocarnivore guild in Midwestern North America. Wildlife Monographs 191(1): 1–61.

Lesmeister, D. B., R. S. Crowhurst, M. E. Gompper, and J. J. Millspaugh. 2013. Landscape ecology of eastern spotted skunks in habitats restored for red-cockaded woodpeckers. Restoration Ecology 21(2): 267–275.

Lesmeister, D. B., M. E. Gompper, and J. J. Millspaugh. 2009. Habitat selection and home range dynamics of eastern spotted skunks in the Ouachita Mountains, Arkansas. The Journal of Wildlife Management 73(1): 18–25.

Taal Levi

DEPT. OF Fisheries and wildlife

I combine empirical data, fieldwork, and quantitative methods to address applied problems in ecology, conservation, and management. My focus is broad, extending from understanding how to assess the spatial extent and ecological consequences of wildlife overexploitation, to fisheries management, the ecology and conservation of predators, disease ecology, and population dynamics in a changing climate. In each case, anthropogenic impacts have altered biotic interactions, and these alterations have consequences for things that humans care about, which is where management options need to be considered. Although my work is rooted in ecology and applied mathematics, I like to cross discipline boundaries to explore implications for human livelihoods and health. 

Current projects include mapping the distribution of rare forest carnivores and determining the causes of their rarity, projecting mountain goat populations under climate change scenarios, understanding the relationship between Amazonian forest fragmentation and the dynamics of infectious disease, understanding the ecology and natural history of semi-aquatic jaguars in the Pantanal of Brazil, understanding the community ecology of brown bears in salmon systems, as well as a variety of projects using environmental DNA to detect fish and quantify their abundance, and several projects using high-throughput sequencing to determine the diets of Alexander Archipelago wolves, fishers, martens, bats, and a variety of large mammals.


Peres,  C.A., Emilio, T., Schietti, T., Desmoulière, S.J.M, Levi, T.  2016. Dispersal limitation induces long-term biomass collapse in overhunted Amazonian forests. Proceedings of the National Academy of Sciences113: 892-897

Levi, T., Massey, A.L., Holt, R.D., Keesing, F., Ostfeld, R.S., Peres, C.A. 2016. Does biodiversity protect humans against infectious disease?: Comment. Ecology. 97: 536-546

Ji, Y., Ashton, L., Pedley, S.M., Edwards, D.P., Tang, Y., Nakamura, A., Kitching, R., Dolman, P.M., Woodcock, P., Edwards, F.A., Larsen, T.H., Hsu, W.W., Benedick, S., Hamer, K.C., Wilcove, D.S., Bruce, C., Wang, X., Levi, T., Lott, M., Emerson, B.C., Yu, D.W. 2013. Reliable, verifiable, and efficient monitoring of biodiversity via metabarcoding. Ecology Letters. 16(10) 1245-1257

Pasari, J.,.Levi, T., Zavaleta, E., Tilman, D. 2013Several scales of biodiversity affect ecosystem multifunctionality. Proceedings of the National Academy of Sciences.110:10219-10222

Levi, T., Kilpatrick, A.M., Mangel, M., Wilmers, C.C. 2012Deer, predators, and the emergence of Lyme disease. Proceedings of the National Academy of Sciences.109:10942-10947

Levi, T., Darimont, C.T., MacDuffee, M., Mangel, M., Paquet, P., Wilmers, C.C. 2012Using Grizzly bears to assess harvest-ecosystem tradeoffs in salmon fisheries. PLoS Biology. 10(4)

Levi, T., Wilmers, C.C. 2012. Wolves-Coyotes-Foxes: A cascade among carnivores. Ecology. 93:921-929

Levi, T., Shepard Jr., G.H., Ohl-Schacherer, J., Peres, C.A., Yu, D.W. 2009. Modeling the long-term sustainability of indigenous hunting in Manu National Park, Peru: Landscape-scale management implications for Amazonia. Journal of Applied Ecology 46: 804-814.

Brenda McComb



Forest structure and composition is modified at stand and landscape scales by natural disturbances and management activities. I study the impacts of these types of natural and human-caused disturbances on vertebrates at stand and landscape scales. Silvicultural and landscape plans can be developed to recover habitat for forest species lost by past management decisions, and should be based on available science, analyzed within an interdisciplinary framework that considers ecological functions, economic realities, and a broad range of social acceptabilities.  I enjoy working within this interdisciplinary framework to ensure that the results of our work are grounded in reality, while also allowing me to learn from experts in other disciplines. Graduate students with whom I have worked have explored a wide range of topics across multiple disciplines, but all of whom had conservation of biodiversity as the practical underpinning to their work.

Selected publications

Kline, J.D., M.E. Harmon, T.A. Spies, A.T. Morzillo, R.J. Pabst, B.C. McComb, F. Schnekenburger, K.A. Olsen, B. Csuti, and J.C. Vogeler (2016) Evaluating carbon storage, timber harvest, and potential habitat possibilities for a western Cascades (US) forest landscape. Ecological Applications 26: 2044-2059.

McComb, B.C. (2015) Wildlife Habitat Management: Concepts and Applications in Forestry, Second Edition. CRC Press, Boca Raton, FL. 401 pp.

McComb, B.C., B. Zuckerberg, D.G. Vesely, and C. A. Jordan (2010) Monitoring Animal Populations and Their Habitats: A Practitioner’s Guide. 277 pp.

Duncan, S.L., B.C. McComb and K. N. Johnson (2010) Integrating ecological and social ranges of variability in conservation of biodiversity: past, present, and future. Ecology and Society 15: 5 [online]

McComb, B. C., T. A. Spies, and K. A. Olsen (2007) Sustaining biodiversity in the Oregon Coast Range: potential effects of forest policies in a multi-ownership province. Ecology and Society 12: 29 [online]

Spies, T.A., K. N. Johnson, K. M. Burnett, J. L. Ohmann, B. C. McComb, G. H. Reeves, P. Bettinger, D. J. Miller, J. D. Kline, and B. Garber-Yonts (2007) Cumulative ecological and socioeconomic effects of forest policies in Coastal Oregon. Ecological Applications 17: 5-17.

Note: Due to a name change, publications prior to 2003 are listed as W. C. McComb

Flather, C. H., K. R. Wilson, D. J. Dean, and W. C. McComb (1997) Identifying gaps in conservation networks: of indicators and uncertainty in geographic‐based analyses. Ecological Applications 7: 531-542.

McGarigal, K. and W. C. McComb (1995) Relationships between landscape structure and breeding birds in the Oregon Coast Range. Ecological Monographs 65: 235-260.

Bruce McCune

Department of Botany and Plant Pathology

I work in three overlapping, interdependent areas:

  • Tools for analysis of ecological communities and habitats: My research on analytical tools concerns how species abundance as a response variable differs from the ideal variables, how this creates problems, how to deal effectively with those problems. My current research in this area focuses on species response surfaces in multidimensional predictor spaces.
  • Ecology of temperate forest epiphytes: My work in this area include basic research on the distribution, abundance, and dispersal limitations of temperate forest epiphytes and applied work on how changing climates and forest practices and will alter our future ecosystems. We have studied the long-term consequences of green-tree retention, how different species and functional groups of lichens are distributed in forests, and how the structure of young forests be changed to enhance the re-entry and development of old-growth associated epiphytes.
  • Taxonomic syntheses of lichens in the Pacific Northwest: My work on regional lichen floras facilitates work on the ecology of lichens in the Pacific Northwest.


McCune, B. 2015. The front door to the fourth corner: variations on the sample unit × trait matrix in community ecology. Community Ecology 16: 267-271.  https://ir.library.oregonstate.edu/xmlui/handle/1957/57993

Smith, R. J., J. C. Benavides, S. Jovan, M. Amacher & B. McCune. 2015. A rapid method for landscape assessment of carbon storage and ecosystem function in moss and lichen ground layers. Bryologist 118: 32-45.  doi: http://dx.doi.org/10.1639/0007-2745-118.1.032

McCune, B. & H. T. Root. 2015. Origin of the dust bunny distribution in ecological community data. Plant Ecology 216: 645-656.

Nelson, P. R., B. McCune, C. Roland & S. Stehn. 2015. Non-parametric methods reveal non-linear functional trait variation of lichens along environmental and fire age gradients. Journal of Vegetation Science 26: 848-865.

Root, H. T., B. McCune & Sarah Jovan. 2014. Lichen communities and species indicate climate thresholds in southeast and south-central Alaska, USA. Bryologist 117: 241-252.

Katie Moriarty

OLYMPIA Forestry Sciences LaboratorY
U.S. Forest Service


My research aims to provide robust science-based information for forest managers and practitioners, specifically aimed at balancing management with the long term conservation of forest-dependent species. We focus on applied hypothesis-based research questions using carnivores, often marten (Martes caurina) and fisher (Pekania pennanti), as study species. We are collecting unprecedented amounts of data relating to species distributions and fine scale habitat use. For instance, my team and I have deployed remote cameras (>2700 locations, >7 million photographs) throughout the forested ranges of Oregon to describe the distribution and detectability of rare coastal marten and fisher, potential prey, and predators. We’re using GPS collars in novel ways to quantify marten and fisher movements, energetics, foraging, and fine-scale selection. Teams are concurrently measuring resting behavior and microsite selection. These multi-scale studies have been possible using an approach, where the support of several small projects funded in part by many institutions – from government agencies to private industry – have combined to produce research where the whole is greater than the sum of its parts.

Selected publications

Moriarty, K. M.*, M.A. Linnell*, B. Chasco, C.W. Epps and W. J. Zielinski (2017) Using high-resolution short-term location data to describe territoriality in Pacific martens. Journal of Mammalogy 98: 679–689. *co-lead authors

Facka, A.N. and K. M. Moriarty (in press) A collaborative research approach for the Martes complex: science for conservation. Accepted chapter.

Moriarty, K. M., J.D. Bailey, S.E. Smythe and J. Verschuyl (2016) Distribution of Pacific marten in coastal Oregon. Northwest Naturalist 97: 71-81.

Moriarty, K. M., C. W. Epps, and W. J. Zielinski (2016) Forest thinning changes movement patterns and habitat use by Pacific marten. Journal of Wildlife Management 80: 621-633.

Moriarty, K. M., C. W. Epps, M. G. Betts, J. Bailey, D. Hance, and W. J. Zielinski (2015) Experimental evidence that simplified forest structure interacts with snow cover to influence functional connectivity for Pacific martens. Landscape Ecology 30: 1865-1877.

Moriarty, K. M., and C. W. Epps (2015) Influence of retained satellite information on global positioning system (GPS) collar performance in a forested ecosystem: an experiment with miniature GPS devices (<60g). Wildlife Society Bulletin 39: 349-357.

Moriarty, K.M., W.J. Zielinski, and E.D. Forsman (2011) Decline in American marten occupancy rates at Sagehen Experimental Forest, California. Journal of Wildlife Management 75: 1774-1787.

Ian Munanura

DEPT. OF forest ecosystems & society


I lead the International Program for Ecotourism, Endangered Species and Family Wellbeing at Oregon State University. Our program aims to identify and communicate the critical family resiliency and wellbeing constraints that may be responsible for ecologically destructive forest-use behavior of rural forest-adjacent communities, and the remedial potential of ecotourism and community engagement. We work with local partners worldwide to conduct applied research in forest landscapes with exceptional concentrations of endemic and endangered wildlife species that are experiencing loss of habitat due to human livelihood activities.  We use the Family Stress Theory to explore family stressing events, and family capability constraints (both physical and emotional) that may be responsible for family wellbeing crisis, resulting in desperate and destructive forest-use behavior to maintain livelihoods. In addition, our research program explores the potential for ecotourism (social entrepreneurship), and community engagement (participatory forest management, managed access to non-timber forest products) to promote integrated conservation of endangered species and human wellbeing. Current research projects are in Rwanda (communities neighboring Mountain Gorilla habitat in Volcanoes National Park), Uganda (communities neighboring Mountain Gorilla habitat in Mgahinga National Park), Indonesia (communities neighboring Orangutan habitat in Gunung Leuser National Park), and Oregon, USA (rural communities within the Northwest Forest Plan region). We are also developing projects in Malaysia (communities neighboring Orangutan habitat in Sabah region of Borneo Island). Future research will expand our focus to include new sites in Madagascar, South America and other regions, where critically endangered wildlife species continue to be threatened by human livelihood activities.

Selected publications

Munanura, I. E., Backman K. F., Hallo, C. J. & Powell, B. R. (2016) Perceptions of tourism revenue sharing impacts on Volcanoes National Park, Rwanda: a Sustainable Livelihoods framework. Journal of Sustainable Tourism 24: 1709-1726.

Backman K. F., & Munanura, I. E. (2015) Introduction to the special issues on ecotourism in Africa over the past 30 years. Journal of Ecotourism 14: 95-99.

Munanura, I. E., Backman K. F., Moore, D. D., Hallo, C. J., & Powell, B. R. (2014) Household poverty dimensions influencing forest dependence at Volcanoes National Park, Rwanda. An application of the sustainable livelihoods framework. Natural Resources 5: 1031-1047.

Munanura, I. E., Backman, K. F & Sabuhoro, E. (2013) Managing tourism growth in endangered species habitats of Africa: Volcanoes National Park in Rwanda. Current Issues in Tourism 16: 700-718.

Mark D. Needham



I am a social scientist who focuses on understanding human experiences and behavior within the context of nature, and using this to inform land management and advance scientific thought.  My work is informed by concepts and theories grounded in social psychology and driven by questions about how people value, experience, and impact recreation and tourism settings (e.g., parks, protected areas) and other natural resources (e.g., wildlife, forests, marine areas).  I also have expertise in survey design and administration, and advanced statistical methods.  My most recent work on forest biodiversity issues includes studies of public attitudes toward using biotechnological (e.g., genetic modification) and non-biotechnological (e.g., tree breeding, assisted migration) interventions to address forest health threats (e.g., diseases such as chestnut blight, pests such as pine beetle, climate change).  I am also working on a study of public tradeoffs of ecosystem services associated with aerial herbicide spraying and other intensive forest management practices on private forestlands. I am Editor-in-Chief of the international journal Human Dimensions of Wildlife, and I am also Director of the Natural Resources, Tourism, and Recreation (NATURE) Studies Lab at Oregon State University.


SELECTED publications

Needham, M. D., Vaske, J. J., & Petit, J. D. (2017). Risk sensitivity and hunter perceptions of chronic wasting disease risk and other hunting, wildlife, and health risks. Human Dimensions of Wildlife, 22(3), 197-216.

Perry, E. E., Needham, M. D., & Cramer, L. A. (2017). Coastal resident trust, similarity, attitudes, and intentions regarding new marine reserves in Oregon. Society and Natural Resources, 30(3), 315-330.

Strauss, S. H., Jones, K. N., Lu, H., Petit, J. D., Klocko, A. L., Betts, M. G., Brosi, B. J., Fletcher Jr., R. J., & Needham, M. D. (2017). Reproductive modification in forest plantations: Impacts on biodiversity and society. New Phytologist, 213(3), 1000-1021.

Morzillo, A. T., & Needham, M. D. (2015). Landowner incentives and normative tolerances for managing beaver impacts. Human Dimensions of Wildlife, 20(6), 514-530.

Needham, M. D., & Vaske, J. J. (2013). Activity substitutability and degree of specialization among deer and elk hunters in multiple states. Leisure Sciences, 35(3), 235-255.

Google Scholar, ResearchGate


Ronald P. Neilson

dept. of botany & Plant pathology

Ronald P. Neilson is a Professor (Courtesy) with the Department of Botany and Plant Pathology and the Department of Forest Science at Oregon State University, recently retired as a BioClimatologist with the USDA Forest Service, Pacific Northwest Research Station. Dr. Neilson has focused on the theory, mechanisms and simulation of vegetation distribution for over three decades. He received the Cooper Award from the Ecological Society of America for his research on oak distribution in the Rocky Mountain region. Dr. Neilson’s MAPSS biogeography model and MC1 dynamic general vegetation model have contributed to national and global assessments by the Intergovernmental Panel on Climate Change (IPCC) and the U.S. Global Change Research Program and to Our Changing Planet, the formal description of the U.S. Global Change Research Program. Dr. Neilson was the lead author for the Forest sector for the IPCC’s special report on The Regional Impacts of Climate Change and the convening lead author for an Annex to the Special Report on simulations of global vegetation re-distribution under climate change. Dr. Neilson’s work with the IPCC contributed to their share of the 2007 Nobel Peace Prize, along with former Vice-President, Al Gore. His current work extends into Earth System Modeling, Landscape System Modeling and large-scale fire forecasting.

SELECTED publications

Neilson, R.P. and S.W. Running. 1996. Global dynamic vegetation modelling: coupling biogeochemistry and biogeography models, p. 451-465. In: B. Walker and W. Steffen (eds.), Global Change and Terrestrial Ecosystems. Cambridge University Press, Cambridge.

VEMAP Members. 1995. Vegetation/ ecosystem modeling and analysis project: Comparing biogeography and biogeochemistry models in a continental-scale study of terrestrial ecosystem responses to climate change and CO2 doubling. Global Biogeochemical Cycles 9:407-37.

Neilson, R.P. and D. Marks. 1994. A global perspective of regional vegetation and hydrologic sensitivities from climatic change. J. Vegetation Sci. 5:715-730.

King, G.A. and R.P. Neilson. 1992. The transient response of vegetation to climate change: A potential source of CO2 to the atmosphere. Water, Air and Soil Pollution 64:365-383.

Michael Paul Nelson


Michael Paul Nelson is an environmental scholar, writer, teacher, speaker, consultant, and Professor of Environmental Ethics and Philosophy. He holds the Ruth H. Spaniol Chair of Renewable Resources and serves as the Lead Principal Investigator for the HJ Andrews Experimental Forest Long-Term Ecological Research program at Oregon State University. He is the philosopher in residence of the Isle Royale Wolf-Moose Project, the longest continuous study of a predator-prey system in the world. Michael is the co-founder and co-director of the Conservation Ethics Group, an award-winning environmental ethics consultancy group fusing ethics with social and ecological science, and serves as a senior fellow for the Spring Creek Project for Ideas, Nature, and the Written Word

He is the author of many articles, and the author or editor of a number of books, in and around the area of environmental ethics. Michael is called upon regularly by various government agencies and conservation organizations to assist with understanding the ethical implications of natural resource management decisions.


SELECTED publications

“For goodness sake! What is intrinsic value and why should we care?” (with Chelsea Batavia) Biological Conservation 209 (2017) pp. 366-376.

“Evaluating the principles of wildlife conservation: a case study of wolf (Canis lupus) hunting in Michigan, United States” (with John A. Vucetich, Jeremy T. Bruskotter, Rolf O. Peterson, and Joseph K. Bump), Journal of Mammology, 2017, 98(1):53-64, 2017.

“Field Philosophy: Experiential Environmental Humanities Learning in Isle Royale National Park” (with Lissy Goralnik), 2017, in press (early on line), Environmental Education Research.

“Conserving the world’s megafauna and biodiversity: the fierce urgency of now” (with William J. Ripple, Guillaume Chapron, José Vicente López-Bao, Sarah M. Durant, David W. Macdonald, Peter A. Lindsey, Elizabeth L. Bennett, Robert L. Beschta, Jeremy T. Bruskotter, Ahimsa Campos-Arceiz, Chris T. Darimont, Richard T. Corlett, Amy J. Dickman, Rodolfo Dirzo, Holly T. Dublin, James A. Estes, Kristoffer T. Everatt, Mauro Galetti, Varun R. Goswami, Matt W. Hayward, Simon Hedges, Luke T. B. Hunter, Graham Kerley, Taal Levi, John C. Morrison, Michael Paul Nelson, Thomas M. Newsome, Luke Painter, Robert M. Pringle, Christopher J. Sandom, John Terborgh, Adrian Treves, Blaire Van Valkenburgh, John A. Vucetich, Aaron J. Wirsing, Arian D. Wallach, Christopher Wolf, Rosie Woodroffe, Hillary Young, Li Zhang), 2017, early on line in BioScience.

“The Logical and Practical Necessity of Ethics in Ecological Forestry: A Reply to Palik and D’Amato 2016” (with Chelsea Batavia), January 2017, Journal of Forestry 115(1): 58-59.

“Conservation social science: Understanding and integrating the human dimensions to improve conservation” (with Nathan Bennett; Robin Roth; Sarah C Klain; Kai Chan; Patrick Christie; Douglas A Clark; Georgina Cullman; Deborah Curran; Trevor J Durbin; Graham Epstein; Alison Greenberg; John Sandlos; Richard Stedman; Tara L Teel; Rebecca Thomas; Diogo Veríssimo; Carina Wyborn), 2017, Biological Conservation 205, 93-108.

“H.J. Andrews Forest Discovery: A Conceptual Framework for Interdisciplinary Interpretation and Empathy Development” (with Goralnik, L., O’Connell, K.B., and Schulze, M.), 2016, Proceedings of the 2015 Symposium for Experiential Education Research (SEER), Portland, OR. pp. 30-34.

“Why the North American Model of Wildlife Conservation is Problematic for Modern Wildlife Management” (with Nils Peterson), Human Dimensions of Wildlife, 2017, 22:1(43-54).

“The Moral Urgency of Action to Protect the World’s Megafauna” (with Kathleen Dean Moore) in BioScience, December 2016, 66(12):1009-1010.

“Saving the world’s terrestrial megafauna” (with William J. Ripple, Guillaume Chapron, José Vicente López-Bao, Sarah M. Durant, David W. Macdonald, Peter A. Lindsey, Elizabeth L. Bennett, Robert L. Beschta, Jeremy T. Bruskotter, Ahimsa Campos-Arceiz, Chris T. Darimont, Richard T. Corlett, Amy J. Dickman, Rodolfo Dirzo, Holly T. Dublin, James A. Estes, Kristoffer T. Everatt, Mauro Galetti, Varun R. Goswami, Matt W. Hayward, Simon Hedges, Luke T. B. Hunter, Graham Kerley, Taal Levi, John C. Morrison, Michael Paul Nelson, Thomas M. Newsome, Luke Painter, Robert M. Pringle, Christopher J. Sandom, John Terborgh, Adrian Treves, Blaire Van Valkenburgh, John A. Vucetich, Aaron J. Wirsing, Arian D. Wallach, Christopher Wolf, Rosie Woodroffe, Hillary Young, Li Zhang), BioScience, 66(10):807-812.

Joseph Northrup

Research Scientist wildlife research & monitoring section
Ontario Ministry of Natural Resources and Forestry


My research is focused on understanding the influence of humans on the spatial ecology of wildlife species. The dominant drivers of global environmental change can be measured spatially, and thus a spatial approach is a natural way to understand species responses. Most of my work uses global positioning system radio collar data to understand how species move through and select habitat in landscapes that have been modified by human activities in some way. My research is applied in nature and I aim to address research questions that have meaningful links to conservation policy and wildlife management.

My current research projects span a broad range of taxa and ecosystems, but all are centered around understanding how human-caused environmental change has impacted the spatial ecology of species. I have active research projects on mule deer in Colorado, polar bears at the southern extent of their global range, black bears in Ontario and Marbled Murrelets in the Pacific Northwest.  

Selected publications

Northrup, J.M., Rivers, J.w., Yang, Z. & Betts, M.G. (2019). Synergystic effects of climate and land-use change influence broad-scale avian population declines. Global Change Biology.

Phalan, B.T., Northrup, J.M., Yang, Z., Deal, R.L., Rousseau, J.S., Spies, T.A. & Betts, M.G. (2019) Impacts of the Northwest Forest Plan on forest composition and bird populations. Proceedings of the National Academy of Sciences, doi.org/10.1073/pnas.1813072116

Bastille‐Rousseau, G., Douglas‐Hamilton, I., Blake, S., Northrup, J.M. & Wittemyer, G. (2018) Applying network theory to animal movements to identify properties of landscape space use. Ecological Applications, 28, 854-864.

Northrup, J.M., Anderson, C.R., Hooten, M.B. & Wittemyer, G. (2016) Movement reveals scale dependence in habitat selection of a large ungulate. Ecological Applications, 26, 2746-2757.

Northrup, J.M., Anderson, C.R. & Wittemyer, G. (2015) Quantifying spatial habitat loss from hydrocarbon development through assessing habitat selection patterns of mule deer. Global Change Biology, 21, 3961-3970.

Northrup, J.M., Anderson, C.R. & Wittemyer, G. (2015) Environmental dynamics and anthropogenic development alter philopatry and space-use in a North American cervid. Diversity and Distributions.

Wittemyer, G., Northrup, J.M., Blanc, J., Douglas-Hamilton, I., Omondi, P. & Burnham, K.P. (2014) Illegal killing for ivory drives global decline in African elephants. Proceedings of the National Academy of Sciences, 111, 13117-13121.

Northrup, J.M. & Wittemyer, G. (2013) Characterising the impacts of emerging energy development on wildlife, with an eye towards mitigation. Ecology letters, 16, 112-125.

Christine Olsen


Research Interests: 

  • Natural Resources

  • Social Science

Social science, human dimensions of natural resources, planning and decision-making processes, citizen-agency communication and interactions, forest management before and after wildfires, coupled human-natural systems

SELECTED publications

Olsen, C.S., Toman, E., and S.S. Frederick. In review. A multi-region analysis of factors that influence public acceptance of smoke from different fire sources. International Journal of Wildland Fire.

Frederick, S.S., Olsen, C.S., and E. Toman. In review. Public perceptions of smoke and agency relationships before and after an active fire season: Longitudinal panel results from northern California. Journal of Forestry.

Olsen, C.S., Kline, J., Ager, A., Olsen, K., and K. Short. 1st Revision. Examining the influence of biophysical conditions on WUI homeowners’ wildfire risk mitigation activities in fire-prone landscapes. Ecology & Society.

Kline, J., White, E.M., Fischer, A.P., Steen-Adams, M.M., Charnley, S., Olsen, C.S., Platt, E.K., Bailey, J.D., and T.A. Spies. 1st Revision. Integrating social science information about actor behavior into empirical models of couple human and natural systems. Ecology & Society.

Engebretson, J.M., Hall, T.E., Blades, J.J., Olsen, C.S., Toman, E., and S. Frederick. In Press. Understanding public tolerance of smoke from wildland fires across the United States. Journal of Forestry.

McGee, T., Curtis, A., McFarlane, B., Shindler, B., Christianson, A., Olsen, C.S., and S. McCaffrey. In Press. Facilitating knowledge transfer between researchers and wildfire practitioners. The Forestry Chronicle.

Fischer, A.P., Spies, T.A., Steelman, T.A., Moseley, C., Johnson, B.R., Bailey, J.D., Ager, A.A., Bourgeron, P., Charnley, S., Collins, B.M., Kline, J.D., Leahy, J.E., Littell, J.S., Millington, J.D.A., Nielsen-Pincus, M., Olsen, C.S., Paveglio, T.B., Roos, C.I., Steen-Adams, M.M., Stevens, F.R., Vukomanovic, J., White, E.M., and D.M.J.S. Bowman. In Press. Wildfire risk as a socio-ecological pathology. Frontiers in Ecology and the Environment.

Olsen, C.S., Mazzotta, D., Toman, E., and A.P. Fischer. 2014. Communicating about smoke from fire: Challenges and ways to address them. Environmental Management 54(3): 571-582.

Spies, T.A., White, E., Kline, J.D., Fischer, P., Agar, A., Bolte, J., Koch, J., Platt, E., Olsen, C.S., Jacobs, D., Shindler, B., Steen-Adams, M.M., and R. Hammer. 2014. Examining fire-prone forest landscapes as coupled human and natural systems. Ecology & Society 19(3): 9. DOI: 10.575/ES-06584-190309

Stidham, M., Olsen, C.S., Toman, E., Frederick, S., McCaffrey, S., and B. Shindler. 2014. Longitudinal social science research in natural resource communities: lessons and considerations. Society and Natural Resources 27(10): 1104-1108.

Olsen, C.S. and E. Sharp. 2013. Building community-agency trust in fire-affected communities in Australia and the United States. International Journal of Wildland Fire 22(6): 822-831.

Davis, E.J., Moseley, C., Olsen, C., Abrams, J., and J. Creighton. 2013. The diversity and dynamism of fire science user needs. Journal of Forestry 111(2): 101-107.

Fischer, A.P., Kline, J., Charnley, S., Olsen, C.S. 2013. Identifying policy target groups with qualitative and quantitative methods: The case of wildfire risk on nonindustrial private forest lands. Forest Policy and Economics 28: 69-78.

Fischer, A.P., Korejwa, A., Koch, J., Spies, T., Olsen, C.S., White, E.M., and D. Jacobs. 2013. Using the Forests, People, Fire agent-based social network model to investigate interactions in social-ecological systems. Practicing Anthropology 35(1): 8-13.

Olsen, C.S., Mallon, A., and B. Shindler. 2012. Public acceptance of disturbance-based forest management: factors influencing support. ISRN Forestry Volume 2012. doi:10.5402/2012/594067

McCaffrey, S. and C.S. Olsen. 2012. Research perspectives on the public and fire management: a synthesis of current social science on 8 essential questions. USDA Forest Service General Technical Report GTR-NRS-104. Newtown Square, PA: Northern Research Station. 40 p. **

Shindler, B., Gordon, R., Brunson, M.W., and C.S. Olsen. 2011. Public perceptions of Sagebrush ecosystem management in the Great Basin. Rangeland Ecology and Management 64(4): 335-347.

Olsen, C. and B. Shindler. 2010. Trust, acceptance, and citizen-agency interactions after large fires: influences on planning processes. International Journal of Wildland Fire 19(1): 137-147.

Toman, E., Shindler, B. and C.S. Olsen. 2008. Communication Strategies for postfire planning: lessons learned from forest communities. In Chavez, D.J., Absher, J.D., and P.L. Winter (eds.). Fire social science research from the Pacific Southwest Station: studies supported by National Fire Plan funds. USDA Forest Service General Technical Report PSW-GTR-209. Albany, CA: Pacific Southwest Research Station. Pages 165-180.

Olsen, C.S. and B.A. Shindler. 2007. Citizen-agency interactions in planning and decisionmaking after large wildfires. USDA Forest Service General Technical Report PNW-GTR-715. Portland, OR: Pacific Northwest Research Station. 37 p.

Deanna H. (Dede) Olson

Pacific Northwest Research Station
U.S. Forest Service


I study the conservation biology, behavioral ecology, and population and community ecology of amphibians and reptiles, and selected other taxa. My current projects address: 1) the effects of forest management practices and riparian buffer widths on headwater habitats and species; 2) microhabitat- to landscape-scale habitat modeling, including the effects of climate change on species and habitats; 3) development of data management and mapping portals for amphibian and reptile diseases; and 4) context matters for amphibian infection by the chytrid fungus Batrachochytrium dendrobatidis.  Recently, I have been working on a variety of syntheses with an amazing group of partners -- for western forests and their management, aquatic invasive species, aquatic biodiversity, and amphibian research and conservation priorities. I also serve in a variety of roles:  Team Leader - Aquatic Ecology and Management team, US Forest Service, PNW; Courtesy faculty, Oregon State University; Associate Editor, Diseases section of Herpetological Review; USA regional lead for the IUCN SSC Amphibian Specialist Group; Bsal Technical Advisory Committee; US federal Northwest Forest Plan Intermediate Management Group for Survey and Manage; and IUFRO Division 8, Aquatic Biodiversity lead.

Selected publications

Olson, D.H. and B. Van Horne (2017) People, Forests, and Change: Lessons from the Pacific Northwest. Island Press, Washington, D.C. 331 pp. Available from Island Press.

Gervasi, S.S., P.R. Stephens, J. Hua, C.L. Searle, J. Urbina, D.H. Olson, B.A. Bancroft, V. Weis, J.I. Hammond and R.A. Relyea. (2017) Linking ecology and epidemiology to understand predictors of multi-host responses to an emerging pathogen, the amphibian chytrid fungus. PLoS ONE 12(1): e0167882.

Grant, E.H.C., E. Muths, R.A. Katz, S. Canessa, M.J. Adams, J.R. Ballard, L. Berger, C.J. Briggs, J.T.H. Coleman, M.J. Gray, M.C. Harris, R.N. Harris, B. Hossack, K.P. Huyvaert, J. Kolby, K.R. Lips, R.E. Lovich, H.I. McCallum, J.R. Mendelson III, P. Nanjappa, D.H. Olson, J.G. Powers, K.L.D. Richgels, R.E. Russell, B.R. Schmidt, A. Spitzen-van der Sluijs, M.K. Watry, D.C. Woodhams, and C.L. White (2017) Using decision analysis to support proactive management of emerging infectious wildlife diseases. Frontiers in Ecology and the Environment 15: 214-221. [abstract]

Leach, J.A., D.H. Olson, P.D. Anderson, and B.N.I Eskelson (2017) Spatial and seasonal variability of forested headwater stream temperatures in western Oregon, USA. Aquatic Sciences 79: 291-307.

Penaluna, B.E., D.H. Olson, R.L. Flitcroft, M. Weber, J.R. Bellmore, S.M. Wondzell, J.B. Dunham, S.L. Johnson, and G.H. Reeves (2016) Aquatic biodiversity in forests: A weak link in ecosystem services resilience. Biodiversity and Conservation doi:10.1007/s10531-016-1148-0

Xie, G.Y., D.H. Olson, and A.R. Blaustein (2016) Projecting the global distribution of the emerging amphibian fungal pathogen, Batrachochytrium dendrobatidis, based on IPCC climate futures. PLoS ONE 11(8): e0160746

Robert Pabst


Program coordinator
Pacific Northwest Permanent Sample Plot Program

MSHpulse_2015-07-30 Rob Pabst on a little water and shade break.jpg

My research focuses on the long-term dynamics of forest processes. I serve as coordinator of the Pacific Northwest Permanent Sample Plot Program (PNW-PSP, http://pnwpsp.forestry.oregonstate.edu/), one of the longest-running research projects of its kind in the world, with tree measurements spanning decades to a century across a network of more than 140 permanent plots in Oregon and Washington. The program’s research objective is to understand the trends, patterns and mechanisms of tree growth, biomass accumulation, tree mortality and tree regeneration, as well as successional changes in forest understories. Many of our permanent plots are stem-mapped, allowing us to examine fine-scale spatial patterns of these processes. I also use simulation models to examine forest successional processes under different forest management regimes.

Selected publications

Acker, S.A., J.A. Kertis and R.J. Pabst. 2017. Tree regeneration, understory development, and biomass dynamics following wildfire in a mountain hemlock (Tsuga mertensiana) forest. Forest Ecology and Management 384: 72-82.

Harmon, M.E. and R.J. Pabst. 2015. Testing hypotheses of forest succession using long-term data: 100 years of observations in the Oregon Cascades. Journal of Vegetation Science 26(4): 722–732.

Pabst, R.J., Goslin, M.N., Garman, S.L., and Spies, T.A. 2008. Calibrating and testing a gap model for simulating forest management in the Oregon Coast Range. Forest Ecology and Management 256:958–972.

Pabst, R.J. and Spies, T.A. 2001. Ten years of vegetation succession on a debris-flow deposit in Oregon. Journal of the American Water Resources Association 37(6): 1693-1708.

Pabst, R.J. and Spies, T.A. 1999. Structure and composition of unmanaged riparian forests in the coastal mountains of Oregon, U.S.A. Canadian Journal of Forest Research 29: 1557-1573.

All available at: https://andrewsforest.oregonstate.edu/publications

Luke Painter



My research has focused primarily on the ecology of animal and plant interactions, particularly trophic cascade effects involving large herbivores and top predators.

Selected publications

W. J. Ripple, Newsome, T. M., Wolf, C., Dirzo, R., Everatt, K. T., Galetti, M., Hayward, M. W., Kerley, G. I. H., Levi, T., Lindsey, P. A., Macdonald, D. W., Malhi, Y., Painter, L. E., Sandom, C. J., Terborgh, J., and Van Valkenburgh, B. Collapse of the world's largest herbivores. Science Advances 1:e1400103 - e1400103. 

L. E. Painter, Beschta, R. L., Larsen, E. J., and Ripple, W. J. Recovering aspen follow changing elk dynamics in Yellowstone: evidence of a trophic cascade?”, Ecology 96:252 - 263.

J. L. Batchelor, Ripple, W. J., Wilson, T. M., and Painter, L. E. Restoration of Riparian Areas Following the Removal of Cattle in the Northwestern Great Basin. Environmental Management 55:930 - 942. 

W. J. Ripple, Painter, L. E., Beschta, R. L., and C. Gates, C. Wolves, Elk, Bison, and Secondary Trophic Cascades in Yellowstone National Park. The Open Ecology Journal 3:31-37.

L. E. Painter, Beschta, R. L., Larsen, E. J., and Ripple, W. J. After long-term decline, are aspen recovering in northern Yellowstone?. Forest Ecology and Management 329:108-117.

L. E. Painter and Ripple, W. J. Effects of bison on willow and cottonwood in northern Yellowstone National Park. Forest Ecology and Management 264:50-158.

L. E. Painter. RESEARCH ARTICLE: Redefining Old-Growth in Forested Wetlands of Western Washington. Environmental Practice 11:68. 


Ben Phalan



As of 2018, I am now based at the Federal University of Bahia, Salvador, Brazil. My research is focused on how to reconcile human demands for food, wood and other products with the conservation of birds and other biodiversity. I am interested in understanding the consequences of different production trajectories for biodiversity, and in evaluating potential conservation strategies. I address these questions through fieldwork, data syntheses and spatial analysis. Collaborations include work with the US Forest Service, BirdLife International, the Royal Society for the Protection of Birds, Rainforest Alliance, the International Institute for Sustainability and the UNEP World Conservation Monitoring Centre. I am also interested more widely in ways of identifying and resolving trade-offs between conservation and development; in the ecology, conservation and restoration of tropical forests, especially in Brazil and West Africa; in finding ways to implement land sparing in practice; in the role of sustainability standards in agriculture; and in defining conservation baselines and objectives.

I am also affiliated with the Cambridge Conservation Initiative (UK) and the International Institute for Sustainability (Brazil).

Selected publications

Benjamin T. Phalan, Joseph M. Northrup, Zhiqiang Yang, Robert L. Deal, Josée S. Rousseau, Thomas A. Spies, Matthew G. Betts (2019) Impacts of the Northwest Forest Plan on forest composition and bird populations. Proceedings of the National Academy of Sciences. DOI:10.1073/pnas.1813072116

Phalan, B., Green, R.E., Dicks, L.V., Dotta, G., Feniuk, C., Lamb, A., Strassburg, B.B.N., Williams, D.R., zu Ermgassen, E.K.H.J. and Balmford, A. (2016) How can higher-yield farming help to spare nature? Science 351: 450–451.

Lamb, A., Balmford, A., Green, R.E. & Phalan, B. (2016) To what extent could edge effects and habitat fragmentation diminish the potential benefits of land sparing? Biological Conservation 195: 264–271.

Phalan, B., Green, R. and Balmford, A. (2014) Closing yield gaps: perils and possibilities for biodiversity conservation. Philosophical Transactions of the Royal Society B 369: 20120285.

Phalan, B., Bertzky, M., Butchart, S. H. M., Donald, P. F., Scharlemann, J. P. W., Stattersfield, A. J. and Balmford, A. (2013) Crop expansion and conservation priorities in tropical countries. PLoS ONE 8: e51759.

Phalan, B., Balmford, A., Green, R. E. and Scharlemann, J. P. W. (2011). Minimising the harm to biodiversity of producing more food globally. Food Policy 36: S62-S71.


Klaus J. Puettmann



Edmund Hayes Professor in Silviculture Alternatives

My recent and future work aims at developing silvicultural treatments that maintain or encourage ecosystem adaptability, while providing income, timber, wildlife habitat, clean water, and other goods at the same time. I work on several experiments that investigate management impacts on vegetation structure and compositions. I am also interested in the question, which spatial scale are most influential on ecosystem processes. For example, under which conditions will landscape level factors override stand level silvicultural treatment or alternatively, under which conditions can small scale silvicultural treatments lead to conditions that override landscape level factors, such as climate change.


K.J. Puettmann, Ares, A, Burton, J.I., and E.K. Dodson. 2016. Forest Restoration Using Variable Density Thinning: Lessons from Douglas-Fir Stands in Western Oregon. Forests 7: 310.

Puettmann, K.J., Wilson, S. McG., Baker, S.C., Donoso, P.J., Droessler, L., Amente, G., Harvey, B.D., Knoke, T., Lu, Y. Nocentini, S., Putz, F.E., Yoshida, T., and Bauhus, J. 2015. Silvicultural alternatives to conventional even-aged forest management - what limits global adoption? Forest Ecosystems 2: 8.

Kuehne, C., Weiskittel, A.R., Puettmann, K.J., and S. Fraver. 2015. Effects of thinning induced changes in structural heterogeneity on growth, ingrowth, and mortality in secondary coastal Douglas-fir forests. Canadian Journal of Forest Research 45: 1448–1461.

Messier, C., Puettmann, K., Chazdon, R., Andersson, K.P., Angers, V.A., Brotons, L., Filotas, E., Tittler, R., Parrott, L., and S.A. Levin. 2015. From management to stewardship: viewing forests as complex adaptive systems in an uncertain world. Conservation Letters 8(5): 368–377.

Burton, J.I., Ganio, L., and K.J.Puettmann. 2014. Multi-scale spatial controls of understory vegetation in Douglas-fir - western hemlock forests of western Oregon, USA. Ecosphere 5: art151.

William J. Ripple

Dept. of Forest Ecosystems & Society OREGON STATE UNIVERSITY


William J. Ripple is a Distinguished Professor of Ecology, best known for his research on terrestrial trophic cascades, particularly the role of the gray wolf  in North America as an apex predator and a keystone species that shapes food webs and landscape structures via “top-down” pressures. Ripple heads the Trophic Cascades Program at Oregon State University, which carries out several research initiatives such as the Aspen Project, the Wolves in Nature Project, and the Range Contractions Project.

Selected publications

Ripple W.J., Wolf C., Newsome T.M., Betts M.G., Ceballos G., Courchamp F., Hayward M.W., Van Valkenburgh B., Wallach A.D., Worm B. 2019. Are we eating the world's megafauna to extinction? Conservation Letters. 2019 e12627. 

Beschta R.L., Ripple W.J. 2019. Can large carnivores change streams via a trophic cascade? Ecohydrology. 12(1):e2048.

Dellinger J.A., Shores C.R., Craig A., Heithaus M.R., Ripple W.J., Wirsing A.J. 2019. Habitat use of sympatric prey suggests divergent anti‑predator responses to recolonizing gray wolves. Oecologia.

Ripple W.J., Miller S.D., Schoen J.W., Rabinowitch S.P.. 2019. Large carnivores under assault in Alaska. PLoS Biol. 17(1):e3000090.

Painter L.E., Beschta R.L., Larsen E.J., Ripple W.J.. 2018. Aspen recruitment in the Yellowstone region linked to reduced herbivory after large carnivore restoration. Ecosphere. 9(8):e02376.

Ripple W.J., Wolf C., Newsome T.M., Hoffmann M., Wirsing A.J., McCauley D.J.. 2018. Both the largest and smallest vertebrates have elevated extinction risk. PNAS. 115(26):E5847-E5848.

van Eeden LM, Eklund A, Miller J.R.B., Lopez-Bao J.V., Chapron G., Cejtin M.R., Crowther M.S., Dickman C.R., Frank J., Krofel M et al.. 2018. Carnivore conservation needs evidence based livestock protection. PLoS Biol. 16(9):e2005577.

Dellinger J.A., Shores C.R., Marsh M., Heithaus M.R., Ripple W.J., Wirsing A.J.. 2018. Impacts of recolonizing gray wolves (Canis lupus) on survival and mortality in two sympatric ungulates. Can. J. Zool.. 96:760–768.

Lundgren E.J., Ramp D., Ripple W.J., Wallach A.D.. 2018. Introduced megafauna are rewilding the Anthropocene. Ecography. 41(6):857-866.

Wallach A.D., Lundgren E.J., Ripple W.J., Ramp D.. 2018. Invisible megafauna. Conservation Biology.

Wolf C., Betts M.G., Levi T., Newsome T.M., Ripple W.J.. 2018. Large species within carnivora are large carnivores. Royal Society Open Science. 5:181228.

van Eeden LM, Crowther M.S., Dickman C.R., Macdonald D.W., Ripple W.J., Ritchie E.G., Newsome T.M.. 2018. Managing conflict between large carnivores and livestock. Conservation Biology. 32(1):26-34.

Peters R., et al. 2018. Nature Divided, Scientists United: US–Mexico Border Wall Threatens Biodiversity and Binational Conservation. Bioscience.

Courchamp F., Jaric I., Albert C, Meinard Y., Ripple W.J., Chapron G.. 2018. The paradoxical extinction of the most charismatic animals. PLoS Biol. 16(4):e2003997.

Ripple W.J., Wolf C., Galetti M., Newsome T.M., Green T.L., Alamgir M, Crist E, Mahmoud M.I., Laurance W.F.. 2018. The Role of Scientists’ Warning in Shifting Policy from Growth to Conservation Economy. Bioscience. 68(4):239-240.

Ripple W.J., Meijaard E., Newsome T.M.. 2018. Saving the World with Satire: A Response to Chapron et al.. Trends in Ecology & Evolution. 33(7):483-484.

Shackelford N, Standish R.J., Ripple W.J., Starzomski B.M.. 2018. Threats to biodiversity from cumulative human impacts in one of North America's last wildlife frontiers. Conservation Biology. 32(3):672-684.

Beschta R.L., Painter L.E., Ripple W.J.. 2018. Trophic cascades at multiple spatial scales shape recovery of young aspen in Yellowstone. Forest Ecology and Management. 413:62-69.

Gray T.N.E., Hughes A.C., Laurance W.F., Long B., Lynam A.J., O'Kelly H., Ripple W.J., Seng T., Scotson L., Wilkinson N.M.. 2018. The wildlife snaring crisis: an insidious and pervasive threat to biodiversity in Southeast Asia. Biodivers Conserv. 27:1031–1037.

Beschta R.L., Ripple W.J.. 2018. Wolf-triggered trophic cascades and stream channel dynamics in Olympic National Park: a comment on East et al.. Earth Surf. Process. Landforms.

Jim Rivers

DEPT. OF Forest ecosystems & Society


I am an animal ecologist with research interests that extend into the fields of animal behavior, physiological ecology, and conservation science. The research questions I pursue are grounded in both basic and applied principles, and nearly all are investigated through empirical field studies of wild populations in forested ecosystems. Some of my current projects include studies that are examining bee community response to wildfire and logging, testing the demographic response of early-successional birds to herbicides, evaluating the impacts of supplemental feeding on the behavior and physiology of songbirds, and quantifying the breeding habitat requirements of the Marbled Murrelet.


Principal Investigator

The OSU College of Forestry initiated a long term, comprehensive study in 2015 to assess and understand murrelet habitat needs in relation to a number of forest management issues. This research applies new methods and technology to improve our understanding of murrelet nesting habitat requirements and factors that affect breeding success.

Selected publications

Rivers, J. W., G. N. Newberry, C. J. Schwarz, and D. R. Ardia. 2016. Success despite the stress: violet-green swallows increase glucocorticoids and maintain reproductive output following experimental increases in flight costs. Functional Ecology 30:xx-xx.

Rivers, J. W., M. Johnson, S. M. Haig, C. J. Schwarz, J. Burnett, J. Brandt, D. George, and J. Grantham. 2014. An analysis of monthly home range size in the critically endangered California Condor. Bird Conservation International 24:492-504.

Rivers, J. W., M. A. Blundell, and S. I. Rothstein. 2014. Mismatched begging displays between foreign and host offspring reduce brood parasite fitness. Behavioral Ecology 25:785-793.

Rivers, J. W., A. L. Liebl, J. C. Owen, L. B. Martin, and M. G. Betts. 2012. Baseline corticosterone is positively related to juvenile survival in a migrant passerine bird. Functional Ecology 26:1127-1134.

Rivers, J. W., G. S. Gipson, D. P. Althoff, and J. S. Pontius. 2010. Long-term community dynamics of small landbirds with and without exposure to extensive disturbance from military training activities. Environmental Management 45:203-216.

Douglas Robinson

DEPT. OF fisheries and wildlife


I currently focus on the influence of environmental change on birds. My students and I value strengthening human perspective regarding the degree to which bird populations and distributions have changed over time. Part of developing that perspective is to help people understand the value of creating high quality benchmarks of where birds live now and how many we have. Future generations will thank us for having the foresight to give them such valuable legacies of information. My work has also involved studies of why communities fall apart after habitat fragmentation, evolution of life histories, and development of tracking devices to follow animals.

Selected publications

See Google Scholar

Nathan Schumaker

Research Ecologist

EPA - Health & Environmental Effects Research Laboratory

Nathan Schumaker.png

I specialize in the development of predictive wildlife models. My research spans the fields of conservation biology, landscape ecology, genetics, and epidemiology. The overarching goal of my work is the development of scientifically defensible theory, technology, and methods that improve wildlife conservation and inform policy. To this end, I collaborate with scientists and conservation practitioners throughout the world. Below, I describe some focal areas that characterize my contributions.

Simulation modeling

  • Designed and developed the HexSim life history simulator.

Conservation practice

  • Used HexSim, and its predecessor PATCH, in the development of spatial simulation models for >55 different wildlife species.

Conservation theory

  • Conceived of and constructed novel research methods that bring biological detail, individual behavior, species interactions, and meaningful disturbance regimes to assessments of population trends, source-sink dynamics, and measures of landscape connectivity. Developed new approaches to predict spatio-temporal patterns of gene flow and disease spread in real-world systems.


Dunk JR, Woodbridge B, Schumaker NH, Glenn EM, White B, LaPlante DW, et al. 2019. Conservation planning for species recovery under the Endangered Species Act: A case study with the Northern Spotted Owl. PLoS ONE 14(1): e0210643. https://doi.org/10.1371/journal. pone.0210643.

Heinrichs JA, Lawler JJ, Schumaker NH, Walker L, Cimprich D, Bleisch A. Assessing source-sink stability in the context of management and land-use change. Landscape Ecology (in press).

Dickson BG, Albano CM, Anantharaman R, Beier P, Fargione J, Graves TA, Gray ME, Hall KR, Lawler JJ, Leonard PB, Littlefield CE, McClure ML, Novembre J, Schloss CA, Schumaker NH, Shah VB, Theobald DM. Circuit-theory applications to connectivity science and conservation. Conservation Biology (in press).

Schumaker NH, Brookes A. 2018. HexSim: a modeling environment for ecology and conservation. Landscape Ecology 33:197-211.

Heinrichs JA, Lawler JJ, Schumaker NH, Wilsey CB, Monroe KC, Aldridge CL. 2018. A multispecies test of source.sink indicators to prioritize habitat for declining populations. Conservation Biology 32:648-659.

Heinrichs JA, Aldridge CL, Gummer DL, Monroe AP, Schumaker NH. 2018. Prioritizing actions for the recovery of endangered species: Emergent insights from Greater Sage-grouse simulation modeling. Biological Conservation 218:134-143.

Pacioni C, Kennedy MS, Berry O, Stephens D, Schumaker NH. 2018. Spatially-explicit model for assessing wild dog control strategies in Western Australia. Ecological Modelling:368:246-256.

Heinrichs JA, Aldridge CL, O’Donnell M, Schumaker NH. 2017. Using dynamic population simulations to extend resource selection analysis and prioritize habitat for conservation. Ecological Modelling 359:449-459.

Wiens JD, Schumaker NH, Inman RD, Esque TC, Longshore KM, Nussear KE. 2017. Spatial demographic models to inform conservation planning of Golden Eagles in renewable energy landscapes. Journal of Raptor Research 51:234-257.

Pierson JC, Coates DJ, Oostermeijer GB, Beissinger SR, Bragg JG, Sunnucks P, Schumaker NH, Young AG. 2016. An assessment of the consideration of genetic factors in threatened species recovery plans on three continents. Frontiers in Ecology and the Environment 14:433-440.

David C. Shaw

Associate ProfessoR, Department of Forest Engineering, Resources, and Management

Forest Health Specialist, Forestry and Natural Resources Extension


I am a forest health specialist and forest biologist who studies applied and basic aspects of forest pathology (fungi), forest entomology (insects), and forest parasitic plants (mistletoe), as well as their interactions with tree mortality, forest disturbance, fire, drought and Silviculture.  I am Director of the Swiss Needle Cast Cooperative, an applied research cooperative focused on a needle disease of Douglas-fir.  I am also Co-PI for the Permanent Plot System, a long-term vegetation monitoring program sponsored by the USFS Pacific Northwest Research Station and NSF/LTER at the USFS H.J. Andrews Experimental Forest, cooperating with OSU. 

Faculty webpage: http://ferm.forestry.oregonstate.edu/facstaff/shaw-david

Swiss Needle Cast Cooperative webpage: http://sncc.forestry.oregonstate.edu/

Pacific Northwest Permanent Sample Plot Program webpage: http://pnwpsp.forestry.oregonstate.edu/

Selected publications

Shaw, D.C. and M.C. Agne.  2017.  Fire and Dwarf Mistletoe (Viscaceae: Arceuthobium species) in Western North America: Contrasting Arceuthobium tsugense and Arceuthobium americanum. Botany 95(3): 231-246, 10.1139/cjb-2016-0245.

Pritchard, K.R., J.C. Hagar, and D.C. Shaw.  2017.  Avian abundance and diversity are associated with Oak Mistletoe (Phoradendron villosum) in Willamette Valley Quercus Woodlands. Botany 95(3): 283-294. 10.1139/cjb-2016-0249.

Shaw, D.C., T. Woolley, R.G. Kelsey, B.A. McPherson, D. Westlind, D.L. Wood, and E.K. Peterson.  2017.  Surface fuels in recent Phytophthora ramorum created gaps and adjacent intact Quercus agrifolia forests, East Bay Regional Parks, California, USA.  Forest Ecology and Management 384: 331-338.

Ritóková, G., D.C. Shaw, G. Filip, A. Kanaskie, J. Browning, and D. Norlander.  2016.  Swiss needle cast in Western Oregon Douglas-fir plantations: 20-year monitoring results. Forests 7: 155. doi:10.3390/f7080155.

Agne, M. C., D.C. Shaw, T.J. Woolley, and M. E. Queijeiro-Bolaños.  2014.  Effects of dwarf mistletoe on stand structure of lodgepole pine forests 21-28 years post-mountain pine beetle mortality in central Oregon. PLoS ONE 9: e107532.

Tom Spies

DEPT. of forest ecosystems & Society

emeritus scientist
PNW research station USDA forest service

Research Interests: Forest ecology, forest succession, stand and landscape structure and dynamics, old growth forest ecology and conservation; overstory-understory relationships; coupled natural and human systems.

Selected publications

Griffiths, Robert P.; Gray, Andrew N.; Spies, Thomas A. 2010. Soil properties in old-growth Douglas-fir gaps in the western Cascade Mountains of Oregon.

Spies, Thomas A.; Miller, Jay D.; Buchanan, Joseph B.; Lehmkuhl, John F.; Franklin, Jerry F.; Healey, Sean P.; Hessburg, Paul F.; Safford, Hugh D.; Cohen, Warren B.; Kennedy, Rebecca S.H.; Knapp, Eric E.; Agee, James K.; Moeur, Melinda. 2010. Underestimating risks to the northern spotted owl in fire-prone forests: response to Hanson et al.

Healey, Sean; Cohen, Warren; Spies, Thomas A.; Moeur, Melinda; Pflugmacher, Dirk; Whitley, M. German; Lefsky, Michael. 2008. The relative impact of harvest and fire upon landscape-level dynamics of older forests: lessons from the Northwest Forest Plan.

Pabst, Robert J.; Goslin, Matthew N.; Garman, Steven L.; Spies, Thomas A. 2008. Calibrating and testing a gap model for simulating forest management in the Oregon Coast Range.

Thompson, Jonathan R.; Johnson, K. Norman; Lennette, Marie; Spies, Thomas A.; Bettinger, Pete. 2006. Historical disturbance regimes as a reference for forest policy. in a multiowner province: a simulation experiment.

Christopher Still



My research is focused on thermal imaging of terrestrial ecosystems, on the role of clouds in the ecological structure and function of forests, on the global biogeography and biogeochemistry of C4 grasses, and on the isotopic composition of atmospheric CO2 and linkages between the carbon and water cycles at a range of spatial and temporal scales.

Selected publications

Greer, B. T., Still, C., Cullinan, G. L., Brooks, J. R., & Meinzer, F. C. 2017. Polyploidy influences plant–environment interactions in quaking aspen (Populus tremuloides Michx.). Tree Physiology, 1-11.

Baguskas, S.A., Still, C.J., Fischer, D.T., D’Antonio, C.M., and J.Y. King. 2016. Coastal fog during summer drought improves the water status of sapling trees more than adult trees in a California pine forest. Oecologia, DOI: 10.1007/s00442-016-3556-y.

Greer, B. Still, C.J., Tague, C., Roberts, D., and G. Howe. 2016. Populations of aspen (Populus tremuloides Michx.) with different evolutionary histories differ in their climate occupancy. Ecology and Evolution, DOI: 10.1002/ece3.2102.

Williams, A.P., Schwartz, R.E., Iacobellis, S., Seager, R., Cook, B.I., Still, C.J., Husak, G. and J. Michaelsen. 2015. Urbanization causes increased cloud-base height and decreased fog in coastal southern California. Geophysical Research Letters, DOI: 10.1002/2015GL063266.

Still, C.J., Pau, S. and E.J. Edwards. 2014. Land surface skin temperature captures thermal environments of C3 and C4 grasses. Global Ecology & Biogeography, DOI:10.1111/geb.12121.

Thomas Stokely

Dept. Forest Ecosystems and Society

Oregon State University


I am interested in the functional aspects of biodiversity, specifically pertaining to species traits, trophic interactions and responses of species assemblages to disturbance. I am most interested in the effects of land management practices and natural disturbances on wildlife-plant interactions and the role of these interactions in structuring biodiversity. My master’s and doctoral research focused on the interacting effects of wild-ungulates (black-tailed deer and Roosevelt elk) and management intensity on early successional plant communities in young plantations of the Oregon Coast Range. To quantify these interactions, we constructed large wildlife exclosures within experimental forest plantations that were treated with varying intensities of silvicultural herbicides or left untreated. For 6 years, we followed plant species and crop-tree responses to herbicides and herbivory, while documented ungulate foraging via camera traps. My research revealed that the effects of herbivory on plant communities and conifer plantations depended on management intensity, with the strongest effects of herbivory being most evident where more intensive herbicide treatments reduced plant diversity and altered forage composition. I am currently a Post-Doctoral Scholar, working to conclude my efforts on the Intensive Forest Management experiment. My overall aim is to contribute to science-based land management and wildlife habitat conservation by applying collaborative ecological research in a large-scale, long-term context. 

Selected publications

Stokely, T. D., J. Verschuyl, J. Hagar, M. G. Betts. 2018. Herbivory and herbicide interact to drive plant community and plantation establishment. Ecological Applications. DOI: https://doi.org/10.1002/eap.1777

Root, H.T., J. Verschuyl, T. D. Stokely, P. Hammond, M. A. Scherr, M.G. Betts. 2016. Plant diversity begets moth diversity in an intensive forest management experiment. Ecological Applications. 27: 134–142

Betts, M.G. J. Verschuyl, J. Giovanini, T. Stokely, A.J. Kroll. 2013. Initial experimental effects of intensive forest management on avian abundance. Forest Ecology and Management. 310: 1036-1044.

Steve Strauss



Research Interests: 

  • Biotechnology

  • Forest Genetics

I have been working in tree and plant biotechnology—teaching, providing outreach to the public, and conducting research—for more than 30 years. I run an active genetic transformation (gene insertion) lab that has produced thousands of transgenic trees, and we have tested many of these in the field. Because of the demand for reliable information on GMO crops and trees by the public, I have given numerous lectures to the public, wrote papers about policy options for improved regulations, and for ~10 years taught an interdisciplinary course on the science and social views of genes and chemicals in agriculture. 

Selected publications

Strauss SH, Jones KN, Lu H, Petit JD, Klocko A, Betts MG, Brosi BJ, Fletcher RJ, Needham MD.  2017.  Reproductive modification in forest plantations: impacts on biodiversity and societyNew Phytologist. 213:1000-1021.

Chang SMahon ELMacKay HARottmann WHStrauss SHPijut PMPowell WACoffey VLu HMansfield SD et al..  2018.  Genetic engineering of trees: progress and new horizonsIn vitro Cell. Develop. Biol. 54:341-376.

Bagley JCHeming NMGutiérrez EEDevisetty UKMock KEEckert AJStrauss SH.  2018.  Genotyping-by-sequencing and ecological niche modeling illuminate phylogeography, admixture, and Pleistocene range dynamics in quaking aspen (Populus tremuloides)PeerJ Preprints. 6:e27162v1

Nagle MLeBoldus JMKlocko A.  2018.  Host‐Induced Gene Silencing for Pest/Pathogen ControlIn: eLS. John Wiley & Sons, Ltd: Chichester.

Nagle MDéjardin APilate GStrauss SH.  2018.  Opportunities for Innovation in Genetic Transformation of Forest TreesFrontiers in Plant Science. 9

Klocko ALu HMagnuson ABrunner AMMa CStrauss SH.  2018.  Phenotypic Expression and Stability in a Large-Scale Field Study of Genetically Engineered Poplars Containing Sexual Containment TransgenesFront. Bioeng. Biotechnol. 6:100

Lu HKlocko ABrunner AMMa CMagnuson AHowe GTAn XStrauss SH.  2018.  RNA interference suppression of AGAMOUS and SEEDSTICK alters floral organ identity and impairs floral organ determinacy, ovule differentiation, and seed-hair development in PopulusNew Phytologist.

Allen Thompson

Associate Professor of Ethics & Environmental Philosophy School of History, Philosophy & Religion OREGON STATE UNIVERSITY


Allen Thompson is an Associate Professor of Ethics and Environmental Philosophy at Oregon State University in Corvallis, Oregon.

His research concerns broadening our conception of environmental virtue and moral responsibility as a part of understanding human excellence in adapting to the Anthropocene. Presently, Thompson is writing a book on the ethics of novel ecosystems (Routledge, under contract) and is working on two invited essays for Oxford University Press about the plight of future generations after our moral failure and what's left for hope in front of unfolding environmental catastrophes.

Thompson is President of the International Society of Environmental Ethics, a past Fellow at the Rachel Carson Center (Munich, Germany), and the OSU The Center for the Humanities. He co-edited The Oxford Handbook of Environmental Ethics (OUP 2017) and Ethical Adaptation to Climate Change (MIT 2012). Thompson also serves as book review editor for the journal Environmental Ethics and was a guest on the nationally-syndicated NPR program, Philosophy Talk, discussing climate ethics.

A list of his publications can be found Here.

David Turner

DEPT. OF Forest ecosystems & Society

David Turner.jpg

David’s interests lie primarily in the carbon cycle, spatially-distributed ecosystem models, and the application of satellite remote sensing to vegetation analysis.

Selected publications

Turner, S.B., Turner, D.P., Gray, A.N., Fellers, W. 2018. An approach to estimating forest biomass change over a coniferous forest landscape based on tree-level analysis from repeated lidar surveys. International Journal of Remote Sensing. https://doi.org/10.1080/01431161.2018.1528401

Jaeger, W.K., Amos, A., Bigelow, D.P., Chang, H., Conklin, D.R., Haggerty, R., Langpap, C., Moore, K., Mote, P., Nolin, A., Plantinga, A.J., Schwartz, C, Tullos, D., Turner, D.P. 2017. Scarcity amid abundance: Water, climate change, and coupled human-natural system models. Proceedings of the National Academy of Science 114:11884-11889.

Turner, D.P., Conklin, D.R., Vache, K.B., Schwartz, C., Nolin, A.W., Chang, H., Watson, E. Bolte, J.P. 2016. Assessing mechanisms of climate change impact on the upland forest water balance of the Willamette River Basin, Oregon. Ecohydrology. (DOI:10.1002/eco.1776).

Turner, D.P., Ritts, W.D., Kennedy, R.E., Gray, A., Yang, Z. 2016. Regional carbon cycle responses to 25 years of variation in climate and disturbance in the US Pacific Northwest. Regional Environmental Change (Online First). Supplementary Materials

Masek, J.G., Hayes, D.J., Hughes, M.J., Healey, S.P., Turner, D.P. 2015. The role of remote sensing in process-scaling studies of managed forest ecosystems. Forest Ecology and Management 355:109-123.

Turner, D.P., Conklin, D.R., Bolte, J.P. 2015. Impacts of projected climate change on forest land cover and land use in the Willamette River Basin, Oregon. Climatic Change 133:335-348.

Turner, D.P., Ritts, W.D., Kennedy, R., Gray, A., Yang, Z. 2015. Effects of harvest, fire, and pest/pathogen disturbances on the West Cascades ecoregion carbon balance. Carbon Balance and Management 10:12

Turner, D.P., Jacobson, A.R., Ritts, W.D., Wang, W.L., Nemani, R.R. 2013. A large proportion of North American terrestrial carbon uptake is offset by emissions from harvested products, river/stream evasion, and biomass burning. Global Change Biology 19:3516-3528.

Hayes, D., Turner, D. 2012. The need for “apples-to-apples” comparisons of carbon dioxide source and sink estimates. Eos 93(41):404-405.

Hayes, D.J., Turner, D.P., Stinson, G., West, T.O., Wei, Y., Heath, L.S., Birdsey, R.A., deJong, B., McGuire, A.D., Kurz, W.A., Jacobson, A.R., McConkey, B.G., Huntzinger, D.N., Pan, Y., Post, W.M., Cook, R.B. 2012. Reconciling estimates of the contemporary North American carbon balance among an inventory-based approach, terrestrial biosphere models, and atmospheric inversions. Global Change Biology 18:1282-1299.

Turner, D.P., Ritts, D., Yang, Z., Kennedy, R.E., Cohen, W.B., Duane, M.V., Law, B.E. 2011. Decadal trends in net ecosystem production and net ecosystem carbon balance for a regional socioecological system. Forest Ecology and Management 262:1318-1325.

Meigs, G.W., Turner, S.P., Ritts, W.D., Yang Z., Law, B.E. 2011. Detection and simulation of heterogeneous fire effects on pyrogenic carbon emissions, tree mortality, and net ecosystem production. Ecosystems 14:758-775.


Richard (Dick) Waring



My early research involved quantifying the environmental distribution of the flora in the species-rich forests of southwestern Oregon. From that my graduate students and I went on to create some of the first process-based predictive models of forest growth and water use. As part of the International Biological Program in the 1970s, I was involved in constructing more complicated ecosystem models, which proved too difficult to apply. In the early 1980s, outbreaks of bark beetles and spruce budworm provided a chance  to conduct field experiments that helped us better understand  the role of insects  in forests and how we might better manage them. Later in the decade, I became interested in doing climate change research sponsored by the National Aeronautic and Space Administration (NASA) Biodiversity and Ecological Forecasting. We started to design models that took advantage of information acquired from earth-orbiting satellites to predict productivity, biodiversity, and the susceptibility of forests to natural agents of disturbance.

Selected publications

Waring, R.H.  1982.  Land of the giant conifers.  Natural History 91:54-63.pdf  

Ryan, M.G., Oren, R., Waring, R.H. 2018. Fruiting and sink competition. Tree Physiology 38:1261-1266. pdf

Mathys, A.S., Coops, N.C., Simard, S.W., Waring, R.H., Aitken, S.N. 2018. Diverging distribution of seedlings and mature trees reflects recent climate change in British Columbia. Ecological Modelling 384:145-153.pdf

Boyle, J.R., Tappeiner, J.C., Waring, R.H., Tattersall Smith, C. 2016. Sustainable Forestry: Ecology and Silviculture for Resilient Forests. Reference Module in Earth Systems and Environmental Sciences, Elsevier, doi:10.1016/B978-0-12-409548-9.09761-x.pdf

Mathys, A.S., Coops, N.C., Waring, R.H. 2016. An ecoregion assessment of projected tree species vulnerabilities in western North America. Global Change Biology,doi: 10.1111/gcb.13440.pdf.

Landsberg, J., Waring, R. 2017. Water relations in tree physiology: where to from here? Tree Physiol. 37:18-32.pdf

 Waring, R.H., Coops, N.C. 2016. Predicting large wildfires across western North America by modeling seasonal variation in soil water balance. Climatic Change 135:325–339. pdf

 Waring, R.H., Gao, L. 2016. Recent reduction in the frequency of frost accounts for most of the increased growth of a high elevation spruce forest in northwestern China. Trees-Structure and Function 30: 1225-1236. pdf

Coops, N.C.,  Waring, R.H., Plowright, A.,  Lee, J., Dilts, T.E. 2016. Using remotely-sensed land cover and distribution modeling to estimate tree species migration in the Pacific Northwest Region of North America. Remote Sensing :8, 65; doi:10.3390/rs8010065.pdf

Law, B.E., and Waring, R.H. 2015. Carbon implications of current and future effects of drought, fire, and management on the Pacific Northwest forests. Forest Ecology & Management 355: 4-14. pdf

 Peterman, W.L., and Waring, R.H. 2014. Overshoot in leaf development of ponderosa pine in wet years leads to bark beetle outbreaks on fine-textured soils in drier years. Forest Ecosystems (on line) .pdf

 Waring, R.H., Coops, N.C., Mathys, A., Hilker, T. and Latta, G. 2014. Process-based modeling to assess the effects of recent climatic variation on site productivity and forest function across western North America. Forests5:518-534.pdf  

Dana Warren



My research explores connections between aquatic and terrestrial environments with a focus on stream ecosystems.  I am particularly interested in understanding how disturbance – and recovery from disturbance – in upland and streamside (riparian) forests affects stream biota and stream ecosystem processes over both short and long time scales.  Anthropocentric clearing of upland and riparian forests was prevalent during the 20th century.  Forests are recovering, but under conditions that differ from those in which they initially grew.  A key goal in my research is to determine how long-term trajectories of change and multiple stand development trajectories in forest regeneration can affect the long-term trajectories of change in associated streams.  This effort includes a wide range of individual research projects encompassing field experiments, stream network analyses, and space-for-time studies.  We quantify processes and biota at scales that range from the stream networks,  to headwater subbasins, to stream reaches, to individual rocks.   Beyond streams, my group also explores processes in lakes and how climate change as well as recovery from historic acid deposition across the landscape affect the lake ecosystem, with a particular focus on coldwater fish.

Selected publications

Warren, D.R., C.E. Kraft, D.C. Josephson, and C.T. Driscoll. In Press.  Acid rain recovery, changing climate, and the future of coldwater fisheries.  Submitted to Global Change Biology

Bechtold, H.A., E.J., Rosi-Marshall, D.R. Warren, and W.S. Keeton. In Press. Forest age influences in-stream ecosystem processes in Northeastern US. Ecosystems

Warren, D.R., W.S. Keeton, P.M. Kiffney, M.J. Kaylor, H.A. Becthold, and J. Magee. 2016. Changing forests – changing streams: Riparian forest stand development and ecosystem function in temperate headwaters. Ecosphere 7(8):e01435.

Warren, D.R., S.M. Collins, E.M Purvis, M.J. Kaylor, and H.A. Becthold. 2016. Spatial variability in light yields co-limitation of primary production by both light and nutrients in a forested stream ecosystem. Ecosystems

Warren, D.R., W. S. Keeton, Bechtold, H.A., E.J. Rosi-Marshall. 2013.  Comparing streambed light availability and canopy cover in streams with old-growth versus early-mature riparian forests in western Oregon. Aquatic Sciences 75: 547-558

Warren, D.R., C.E. Kraft, W.S. Keeton, J.S. Nunery, G.E. Likens. 2009. Dynamics of wood recruitment in streams of the northeastern U.S. Forest Ecology and Management 258:804-813

Banner photograph by Matt Kaylor

Todd M. Wilson

Wildlife Biologist
PNW research station
U.S. Forest Service

Todd Wilson.jpg

Todd’s research focuses on:

Effects of forest management activities on spotted owl prey, the ecology of flying squirrels, woodrats, and bats

Effects of forest, riparian, and grassland management activities on small mammals, arboreal rodents, and terrestrial amphibians

Monitoring long-term ecological change using Research Natural Areas

Selected publications

Willson, Todd M., Ph.D..; Forsman, Eric D. 2013. Thinning effects on spotted owl prey and other forest-dwelling small mammals.

Wilson, Todd M., Ph.D..; Schuller, Reid; Holmes, Russ; Pavola, Curt; Fimbel, Robert A.; McCain, Cynthia N.; Gamon, John G.; Speaks, Pene; Seevers, Joan I.; DeMeo, Thomas E.; Gibbons, Steve. 2009. Interagency strategy for the Pacific Northwest Natural Areas Network.

Carey, A.B.; Wilson, T.M., Ph.D..; Maguire, C.C.; Biswell, B.L. 1997. Dens of northern flying squirrels in the Pacific Northwest..

Villa, Lisa J.; Carey, Andrew B.; Wilson, Todd M., Ph.D..; Glos, Karma E. 1999. Maturation and reproduction of northern flying squirrels in Pacific Northwest forests..

Carey, Andrew B.; Maguire, Christine C.; Biswell, Brian L.; Wilson, Todd M., Ph.D. 1999. Distribution and abundance of Neotoma in western Oregon and Washington..

Wilson, T.M., Ph.D..; Carey, A.B. 1996. Observations of weasels in second-growth Douglas-fir forests in the Puget Trough, Washington..

Bayrakci, R.; Carey, A.B.; Wilson, T.M., Ph.D. 2001. Current status of the western gray squirrel population in the Puget Trough, Washington..

Carey, A.B.; Thysell, D.R.; Villa, L.J.; Wilson, T.M.; Wilson, S.M., Ph.D..; Trappe, J.M.; Colgan, W., III.; Ingham, E.R.; Holmes, M. 1996. Foundations of biodiversity in managed Douglas-fir forests..

Batchelor, Jonathan L.; Ripple, William J.; Wilson, Todd M.; Painter, Luke E. 2015. Restoration of riparian areas following the removal of cattle in the northwestern great basin.

Massie, Margaret H.; Wilson, Todd M.; Morzillo, Anita T.; Henderson, Emilie B. 2016. Natural areas as a basis for assessing ecosystem vulnerability to climate change.

Lind, A.J.; Welsh, Hartwell H., Jr..; Wilson, R.A., Ph.D. 1996. The effects of a dam on breeding habitat and egg survival of the foothill yellow-legged frog (Rana boylii).

David Wrathall

College of Earth, Ocean and Atmospheric Sciences


I am part of a team that identified the international cocaine trade as a new major driver of deforestation in Central America. Since the emergence of Central America as a primary transit corridor for the drug trade (in the mid-2000s), over a million hectares of rainforest have been cleared for cattle ranches and other land-intensive activities in spatiotemporal patterns that we can statistically associate with narco-trafficking, and distinguish from background deforestation patterns. Most of this "narco-deforestation" has occurred in biodiversity hotspots, and accounts for 30 to 60% of deforestation occurring within official protected areas --national parks and UN World Heritage Sites comprising the Mesoamerican Biological Corridor-- threatening biodiversity, forest conservation, carbon sequestration, ecological services, and rural and indigenous livelihoods. The influx of drugs into the region is a product of an intensification of the war on drugs pushing traffickers into lower profile transit routes. Deforestation results as traffickers select land-intensive strategies, such as cattle ranching, for laundering the volumes of illicit cash that have poured into the region in recent years. This research has been published in ScienceEnvironmental Research Letters, and the Journal of Latin American Geographyand has been the subject of reporting in the GuardianBBCthe New York TimesNational GeographicScience Now, among many others. I have studied this topic as a principal investigator with grants from the Open Society Foundations, the National Socio-Environmental Synthesis Center (SESYNC), and Future Earth's Program for Early-stage Grants Advancing Sustainability Science (PEGASuS).

I am an assistant professor of geography in the College of Earth, Ocean and Atmospheric Sciences at Oregon State University, where I primarily study climate change and human migration. 

Selected publications

McSweeney, K., Nielsen, E.A., Taylor, M.J., Wrathall, D.J., Pearson, Z., Wang, O., & Plumb, S.T. (2014) Drug policy as conservation policy: narco-deforestation. Science 343: 489-490.

Sesnie, S.E., Tellman, B., Wrathall, D., McSweeney, K., Nielsen, E., Benessaiah, K., ... & Rey, L. (2017) A spatio-temporal analysis of forest loss related to cocaine trafficking in Central America. Environmental Research Letters 12: 054015.

McSweeney, K., Richani, N., Pearson, Z., Devine, J., & Wrathall, D.J. (2017) Why do narcos invest in rural land? Journal of Latin American Geography 16: 3-29.

Zhiqiang Yang

Research Associate
DEPT. of Forest Ecosystems & Society
Oregon state university

Laboratory for applications of remote sensing in ecology (Larse)


I am interested in ecological modeling with remote sensing for landscape and regional ecological information; forest succession; ecological informatics and application development for ecological researches. 

My current work includes linking time series of spectral data with different successional trajectories; spatial modeling of ecosystem production through BiomBGC modeling; developing methods to quantify uncertainties associated with remote sensing analysis.

Selected publications

Cohen WB, Yang Z, Healey SP, Kennedy RE, Gorelick N. 2018. A LandTrendr Multispectral Ensemble for Forest Disturbance Detection. Remote Sensing of Environment 205:131-140.

Healey SP, Cohen WB, Yang Z, Brewer CK, Brooks EB, Gorelick N, Hernandez AJ, Huang C, Hughes MJ, Kennedy RE et al. 2018. Mapping Forest Change using Stacked Generalization: An Ensemble Approach. Remote Sensing of Environment 204:717-728.

King DT, Wang G, Yang Z, Fischer JW. 2017. Advances and Environmental Conditions of Spring Migration Phenology of American White Pelicans. Scientific Reports. 7. 

Davis RJ, Yang Z, Yost A, Belongie C, Cohen WB. 2017. The Normal Fire Environment—Modeling environmental suitability for large forest wildfires using past, present, and future climate normals. Forest Ecology and Management.. 390:173-186.

Cohen WB, Yang Z, Stehman SV, Schroeder TA, Bell DM, Masek JG, Huang C, Meigs GW. 2016. Forest disturbance across the conterminous United States from 1985–2012: The emerging dominance of forest decline. Forest Ecology and Management 360:242-252.

Harold Zald

DEPT. Forestry & wildland resources
Humboldt state university

Harold Zald.jpg

The world’s forests are at a critical moment, where climate change and societies need for ecosystem services (i.e. wood products, carbon sequestration, etc.) are increasing both the stressors on forests, and the value of these ecosystems. My research in the HSU Forest Measurements & Ecology Lab is broadly focused on conducting applied research to develop forest inventory and monitoring methods, assist forest management decision-making, and inform broader forest policy. Within this broader objective, I use field measurements, tree-rings, and remotely sensed imagery across five major research themes:

  • Integrating field and remotely sensed data to generate spatial predictions of forest composition, structure, and change.

  • Ecology and management of Sierra Nevada mixed-conifer forests with emphasis on responses to climate change, alterations of natural disturbance regimes, and ecosystem restoration

  • Subalpine forest-meadow ecotone responses to climate change

  • Responses of tall old trees to long-term climate change and short-term drought events

  • The role of complex terrain and diverse ecosystem configurations in mediating resilience of forests to disturbance agents and climate stressors.

Selected publications

Goodwin, MJ, North MP, Zald HSJ, Hurteau MD. (accepted). The 15-year post-treatment response of a mixed-conifer understory plant community to thinning and burning treatments. Forest Ecology and Management.

Zald HSJ, Dunn CD. (2018). Disentangling drivers of wildfire severity in a multi-owner forest landscape, Oregon USA. Ecological Applications 28: 1068-1080.

Matasci G, Hermosilla T, Wulder MA, White JC, Hobart GW, Coops NC, Zald HSJ. (2018). Large-area mapping of forest structural attributes across the boreal forest of Canada using Landsat pixel composites and lidar plots. Remote Sensing of Environment 209: 90-116.

Matasci G, Hermosilla T, Wulder MA, White JC, Hobart GW, Zald HSJ, Coops NC. (2017). A space-time data cube: Multi-temporal forest structure maps from Landsat and LiDAR. Geoscience and Remote Sensing Symposium (IGARSS), 2017 IEEE International. doi: 10.1109/IGARSS.2017.8127523.

Meigs GW, Zald HSJ, Campbell JL, Keeton WS, Kennedy RE (2016). Do insect outbreaks increase forest fire severity? Environmental Research Letters. doi:10.1088/1748-9326/11/4/045008.

Zald HSJ, Spies TA, Siedl R, Pabst R, Steele EA, Olsen K. (2016). Complex mountain terrain and disturbance history drive patterns of forest carbon density in the western Oregon Cascades, USA. Forest Ecology and Management 366:193-207.

Zald HSJ, Spies TA, Twery M, Harmon ME (2016). Review of forest carbon calculators for managers, policy makers, and educators. Journal of Forestry 114:134-143.

Zald HSJ, Wulder MA, White JC, Hilker T, Hermosilla T, Hobart GW, Coops NC. (2016). Integrating Landsat pixel composites and change metrics with lidar plots to predictively map forest structure and aboveground biomass in Saskatchewan, Canada. Remote Sensing of Environment 176:188-201.

Meigs GW, Campbell JL, Zald HSJ, Kennedy RE, Bailey JD, Shaw DC (2015). Does wildfire likelihood increase following insect outbreaks in conifer forests? Ecosphere 6:art118.

Zald HSJ, Ohmann JL, Roberts HM, Henderson EB, Gregory MJ, McGaughey R, Braaten J (2014) Influence of LiDAR, Landsat time series disturbance history, and plot positional accuracy on the accuracy of imputation maps of forest composition and structure. Remote Sensing of Environment 143:26-38.

Henderson EB, Ohmann JL, Gregory MJ, Roberts HM, Zald HSJ (2014) Species distribution modeling for plant communities: Stacked single-species, or multivariate modeling approaches? Applied Vegetation Science 17:516-527.

Torresan C, Strunk J, Zald HSJ, Zhiqiang Y, Cohen WB (2014) Comparing statistical techniques to classify the structure of mountain forest stands using CHM-derived metrics in Trento province (Italy). European Journal of Remote Sensing 47:75-94.

Zald HSJ, Spies TA, Huso M, Gatziolis D (2012) Climatic, landform, microtopographic, and overstory canopy controls of tree invasion in a subalpine meadow landscape, Oregon Cascades, USA. Landscape Ecology 27:1197-1212.

Zald HSJ (2009) Extent and spatial patterns of grass bald meadow loss (1948- 2000), Oregon Coast Range, USA. Plant Ecology 201:517-529.

Zald HSJ, Gray AN, North M, Kern RA (2008) Initial tree regeneration responses to fire and thinning treatments in a Sierra Nevada mixed-conifer forest, USA. Forest Ecology and Management 256:168-179.

Meyer M, North M, Gray AN, Zald HSJ (2007) Influence of soil thickness on stand characteristics in a Sierra Nevada mixed-conifer forest. Plant and Soil 294:113-123.

Hurteau M, North M, Zald HSJ (2007) Species-specific response to climate reconstruction in upper-elevation mixed-conifer forests of the western Sierra Nevada, California, USA. Canadian Journal of Forest Research 37:1681-1691.

North M, Innes J, Zald HSJ (2007) Comparison of thinning and prescribed fire restoration treatments to Sierran mixed-conifer historic conditions. Canadian Journal of Forest Research 37:331-342.

Gray AN, Zald HSJ, Kern RA, North M (2005) Stand conditions associated with tree regeneration in Sierran mixed-conifer forests. Forest Science 51:198-210.

Wilking M, Juday GP, Barber VA, Zald HSJ (2004) Recent climate warming forces contrasting growth responses of white spruce at treeline in Alaska through temperature thresholds. Global Change Biology 10:1724-1736.

Diego Zárrate-Charry


Research coordinator
ProCAT-Colombia/Sierra to Sea Institute


My experience has been focused in modeling species distributions as a means to conduct conservation planning in tropical ecosystems in Central and South America. I have been part of several biology conservation projects in Colombia, Costa Rica, Peru, and Ecuador using flagship species to identify priority conservation areas. I have also worked on assessment and valuation of environmental services to increase the capacity and management strategies for National Parks agencies in tropical countries.

My primary research interest is to develop species and ecosystem conservation strategies that include and acknowledge regional social and economic realities in tropical countries. Most of my current work is in my native country, Colombia, where the main current conservation threats are landscape transformation and fragmentation, so the issues of habitat conservation and connectivity are critically important.

Selected publications

Zárrate Charry DA, Guerrero-Ochoa I, Jiménez-Alvarado JS, Massey A, Calderon M, Hurtado-Moreno AP, Proeto J, Aconcha Abril, I, Vela Vargas IM, Gonzalez Maya JF (2016) Strategies for human-jaguar conflict resolution in agricultural areas of Sierra Nevada de Santa Marta, Colombia. Wildfelid Monitor 9: 2.

Aconcha-Abril I, Jiménez-Alvarado JS, Moreno-Diaz C, Zárrate-Charry DA & González-Maya JF (2016) Estado del conocimiento del conflicto por grandes felinos y comunidades rurales en Colombia: avances y vacíos de información. Mammalogy Notes 3(1): 46-51.

Jiménez-Alvarado S, Arias-Ocampo A, Pineda-Guerrero A, Zárrate-Charry DA, Vela-Vargas M, Chacón-Pacheco J y González Maya JF. (2016) Análisis de la distribución del grisón (Galictis vittata) (Carnivora: Mustelidae) en el Caribe colombiano. Therya 7 (1): 179-186.

Cruz Rodriguez C, Gonzalez Maya JF, Rodriguez Bolaños A, Cepeda Mercado AA, Zárrate Charry DA, Belant Jorrold L. (2015) Ocelot Leopardus pardalis (Carnivora: Felidae) spatial ecology in a fragmented landscape of Colombia. Revista Mexicana de Mastozoología 5(1): 17-24.

Zárrate Charry DA, Suarez C y Ortega C. (2013) Análisis de bienes y servicios ecosistémicos y aplicación de la herramienta InVEST en El Corredor Trinacional La Paya, Cuyabeno Y Güeppí. Technical report World Wide Fund for Nature WWF. Bogotá, Colombia. 53 pp.

Castaño-Uribe C, Gónzalez-Maya JF, Zárrate Charry DA, Ange-Jaramillo C y Vela-Vargas IM (Eds) (2013) Plan de Conservación de Felinos del Caribe Colombiano: Los felinos y su papel en la planificación regional integral basada en especies clave. Fundación Herencia Ambiental Caribe, ProCAT Colombia, The Sierra to Sea Institute. Santa Marta Colombia.

González-Maya JF, Cepeda AA, Belant JL, Zárrate-Charry DA, Balaguera-Reina S & Rodríguez Bolaños A (2011) Research priorities for the small carnivores of Colombia. Small Carnivore Conservation 44: 7–13.